Dzharaonyx

Dzharaonyx; Temporal range: Late Cretaceous,
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Clade:	Saurischia
Clade:	Theropoda
Family:	†Alvarezsauridae
Subfamily:	†Parvicursorinae
Genus:	†Dzharaonyx; Averianov & Sues, 2022
Species:	†D. eski
Binomial name
	†Dzharaonyx eski; Averianov & Sues, 2022

Dzharaonyx (meaning "Dzharakuduk claw", named after the type locality) is a genus of alvarezsaurid theropod dinosaur from the Late Cretaceous Bissekty Formation of Uzbekistan. The type species is Dzharaonyx eski; eski being an Uzbek word for "old".[1]

Description

Dzharaonyx is known from a series of disassociated but well-preserved postcranial material. The form of the humerus is intermediate between that of Patagonykus and Mononykus.[1]

Classification

Phylogenetic analysis places Dzharaonyx in a polytomy including other Asian members of Parvicursorinae. This makes it the oldest parvicursorine known to date.[1]

Alvarezsauridae

Alvarezsaurus

Achillesaurus

Albertonykus

	Qiupanykus

	Nemegtonykus

Parvicursorinae

Albinykus

Xixianykus

PIN 5838/1 (=Ondogurvel[2])

Khulsanurus

Dzharaonyx

	Mononykus

	Shuvuuia

Linhenykus

	Ceratonykus

	Parvicursor

Paleoenvironment

Dzharaonyx lived in the Bissekty Formation, which has been intensively studied in recent years. It represents a brackish environment.[3] It coexisted with larger theropods including the dromaeosaurid Itemirus, the tyrannosauroid Timurlengia, and the troodontid Urbacodon. The hadrosaur Levnesovia, ankylosaurs, and the sauropod Dzharatitanis represented the herbivores in this ecosystem.

References

Averianov AO, Sues HD (2022). "New material and diagnosis of a new taxon of alvarezsaurid (Dinosauria, Theropoda) from the Upper Cretaceous Bissekty Formation of Uzbekistan". Journal of Vertebrate Paleontology. 41 (5): e2036174. doi:10.1080/02724634.2021.2036174. S2CID 247391327.
Averianov AO, Lopatin AV (2022). "A new alvarezsaurid theropod dinosaur from the Upper Cretaceous of Gobi Desert, Mongolia". Cretaceous Research. 134: 105168. doi:10.1016/j.cretres.2022.105168. S2CID 247000540.
Redman, C.M., and L.R. Leighton. 2009. Multivariate faunal analysis of the Turonian Bissekty Formation: Variation in the degree of marine influence in temporally and spatially averaged fossil assemblages. PALAIOS 24(1). 18–26. Accessed 2019-03-22.

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[Dzharaonyx-1] Averianov AO, Sues HD (2022). "New material and diagnosis of a new taxon of alvarezsaurid (Dinosauria, Theropoda) from the Upper Cretaceous Bissekty Formation of Uzbekistan". Journal of Vertebrate Paleontology. 41 (5): e2036174. doi:10.1080/02724634.2021.2036174. S2CID 247391327.

[2] Averianov AO, Lopatin AV (2022). "A new alvarezsaurid theropod dinosaur from the Upper Cretaceous of Gobi Desert, Mongolia". Cretaceous Research. 134: 105168. doi:10.1016/j.cretres.2022.105168. S2CID 247000540.

[3] Redman, C.M., and L.R. Leighton. 2009. Multivariate faunal analysis of the Turonian Bissekty Formation: Variation in the degree of marine influence in temporally and spatially averaged fossil assemblages. PALAIOS 24(1). 18–26. Accessed 2019-03-22.