Dennstaedtia

Dennstaedtia punctilobula hay-scented fern Hay-scented fern (Dennstaedtia punctilobula) is often considered to be an interfering species in New York forest understories, where large deer populations over-browse other native plants but avoid this fern, which prevents tree regeneration.

Polypodiales — Dennstaedtiaceae — Dennstaedtia
Dennstaedtia	Hay-scented fern	N.Y. Status	Images	Distribution	NY NPT
Dennstaedtia punctilobula (Michx.) T.Moore 1803. Nephrodium punctilobulum Michx. 1810. Aspidium punctilobum Willd. 1827. Sitobolium punctilobum Desv. 1848. Dicksonia punctilobula A.Gray 1857. Dennstaedtia punctilobula T.Moore	Hay-scented fern, Eastern hay-scented fern, Hay-scent fern, Boulder fern Dennstaedtie à   lobules ponctués	Native, C:4, Secure, Interfering FACU-UPL Perennial, Herb-fern			NYFA-5 USDA-NN NPT Can ARS ITIS FNA Trop. Images, wsp

Pteridium

Pteridium aquilinum eastern bracken fern

Polypodiales — Dennstaedtiaceae — Pteridium
Pteridium	Brackenfern	N.Y. Status	Images	Distribution	NY NPT
Pteridium aquilinum  (L.) Kuhn var. latiusculum (Desv.) Underw. ex A. Heller 1827. Pteris latiuscula Desv. 1909. Pteridium aquilinum var. latiusculum 1914. Pteridium latiusculum (Desv.) Hieron. 1941. Pteridium aquilinum ssp. latiusculum	Eastern brackenfern, Eastern bracken, Brackenfern Fougère-aigle de l'Est	Native, C:3, Secure, Interfering FACU Perennial, Fern			NYFA-5 USDA-NN NPT Can ARS ITIS FNA Trop. BNA Images, wsp
Pteridium aquilinum  (L.) Kuhn var. pseudocaudatum (Clute) A. Heller 1900. Pteris aquilina var. pseudocaudata Clute 1900. Pterid. aquilinum var. pseudocaudatum 1919. Pterid. latiusculum var. pseudocaudatum 1941. Pterid. aquilinum ssp. pseudocaudatum	Southeastern Bracken, Tailed bracken, Brackenfern, Western brackenfern	Native, C:3, Endangered, DEC-A[1] FACU Perennial, Fern			NYFA-Z USDA-N0 NPT  ITIS FNA Trop. BNA Images, wsp
Pteridium (excluded)	Brackenfern	N.Y. Status	Images	Distribution	NY NPT
Pteridium aquilinum  (L.) Kuhn ssp. aquilinum 1753. Pteris aquilina L. 1799. Asplenium aquilinum (L.) Bernh. 1879. Pteridium aquilinum (L.) Kuhn 1941. Pteridium aquilinum ssp. typicum	Western brackenfern	Native, N.Y. excluded			NYFA-XCLD USDA-NN ARS  FNA Trop. BNA Images, wsp
↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered.

Subfamily Cryptogrammoideae

Cryptogramma

Cryptogramma stelleri slender cliffbrake, Steller’s cliffbrake

Polypodiales — Pteridaceae — Cryptogrammoideae — Cryptogramma
Cryptogramma	Rock-brake	N.Y. Status	Images	Distribution	NY NPT
Cryptogramma stelleri (S.G.Gmel.) Prantl 1768. Pteris stelleri S.G.Gmel. 1882. Cryptogramma stelleri Prantl	Fragile rockbrake, Slender cliff brake, Slender rock-brake, Steller's rock brake Cryptogramme de steller	Native, C:9, Vulnerable, DEC-D[1] FACU Perennial, Herb-fern, 2n=60			NYFA-3-4 USDA-N NPT Can ARS ITIS FNA Trop. NSE  IPN  Images, wsp
↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.

Subfamily Pteridoideae

Pteris

Pteris multifida spider brake Pteris multifida, the spider brake was listed as present in New York by the 1955 American Fern Journal, which cited Willard Nelson Clute: New York: It is fond of growing on old walls as far north as New York City — Clute (1938, p. 102).[1] USDA-NRCS Plants Database lists the 1958 American Fern Journal as the source for the plant being listed in New York, but it doesn't appear to be listed in that volume.[2] Pteris multifida is now been listed as "excluded" in the New York Flora Atlas. ↑ G.N. Jones (1955) "On the Occurrence of Pteris multifida in the United States." American Fern Journal, 45:107-111. ↑ USDA-NRCS Plants Database for Pteris multifida (July 21, 2015)

Polypodiales — Pteridaceae — Pteridoideae — Pteris
Pteris (excluded taxa)	Brake-fern	N.Y. Status	Images	Distribution	NY NPT
Pteris multifida  Poir. 1804. Pteris multifida Poir. 1938. Pycnodoria multifida (Poir.) Small  auct.. Pteris serrulata non L.f.	Spider brake, Spider fern, Chinese brake, Huguenot fern, Saw-leaved bracken	Introduced from  temperate Asia,  tropical Asia, N.Y. excluded			NYFA-XCLD USDA-X0 ARS ITIS FNA Trop. IPN  Images, wsp

Subfamily Cheilanthoideae

Pellaea

Polypodiales — Pteridaceae — Cheilanthoideae — Pellaea
Pellaea	Cliff-brake	N.Y. Status	Images	Distribution	NY NPT
Pellaea atropurpurea  (L.) Link Pteris atropurpurea	Purple cliffbrake, Purple-stemmed cliff-brake	Native, Vulnerable, DEC-D[1]			NYFA-4 USDA-NN NPT  Images, wsp
Pellaea glabella  Mett. ex Kuhn ssp. glabella 1858. Pellaea rigida var. rigida 1869. Pellaea glabella var. glabella     Pellaea glabella var. bushii     Pellaea glabella ssp. glabella 1902. Pellaea atropurpurea var. bushii 1938. Pellaea atropurpurea fo. glabella	Smooth cliff-brake Slender cliff-brake Dwarf cliffbrake Simple cliffbrake Pelléade glabre Pelléade glabre typique Pelléade glabre variété glabre	Native, Threatened, DEC-B[2]			NYFA-2 USDA-NN NPT Can FNA Trop. Images, wsp
↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (b) Threatened.

Myriopteris

Polypodiales — Pteridaceae — Cheilanthoideae — Myriopteris
Myriopteris	Lip-fern	N.Y. Status	Images	Distribution	NY NPT
Myriopteris lanosa (Michx.) Grusz & Windham 1803. Nephrodium lanosum Michx. 1804. Adiantum vestitum Spreng. 1806. Cheilanthes vestita Sw. 1859. Cheilanthes lanosa (Michx.) D.C.Eaton 2013. M. lanosa (Michx.) Grusz & Windham	Hairy lipfern, Wooly lip-fern	Native, Endangered, DEC-A[1]			NYFA-Z USDA-N0 NPT  ARS ITIS FNA Trop. BNA Images, wsp
↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered.

Subfamily Vittarioideae

Adiantum

Adiantum pedatum maidenhair fern

Polypodiales — Pteridaceae — Vittarioideae — Adiantum
Adiantum	Maidenhair-fern	N.Y. Status	Images	Distribution	NY NPT
Adiantum pedatum  L. 1753. Adiantum pedatum L.	Maidenhair fern, Northern maidenhair-fern, Northern maidenhair, Five-finger fern Adiante du Canada	Native, Likely secure, DEC-D[1]			NYFA-4 USDA-NN NPT Can ARS  FNA Trop. Images, wsp
Adiantum capillus-veneris  L. 1753. Adiantum capillus-veneris	Venus’s hair fern Common maidenhair Southern maidenhair	Introduced from  southern US, N. America native, Not naturalized			NYFA-Xm USDA-NN NEW-0 ARS  Trop. Images, wsp
↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.

Vittaria

Vittaria appalachiana Appalachian shoestring fern The genus Vittaria is represented in New York by the endangered Appalachian shoestring fern, which has been found in only a few counties and only in its gametophyte stage.[1] ↑ New York Flora Atlas: Vittaria appalachiana appears only as "minute moss-like gametophytes" in New York.

Polypodiales — Pteridaceae — Vittarioideae — Vittaria
Vittaria	Shoestring-fern	N.Y. Status	Images	Distribution	NY NPT
Vittaria appalachiana Farrar & Mickel (1991)	Appalachian shoestring fern, Appalachian vittaria, Appalachian gametophyte fern	Native, Endangered, DEC-A[1]			NYFA-1 USDA-N0 Images, wsp
↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered.

Subfamily Cystopteridoideae

The Cystopteridoideae Ching & Z.R.Wang (1997) (bladder-ferns)...

Cystopteris

Cystopteris bulbifera bulblet fern

Polypodiales — Aspleniaceae — Cystopteridoideae — Cystopteris
Cystopteris	Bladder-fern	N.Y. Status	Images	Distribution	NY NPT
Cystopteris bulbifera  (L.) Bernh. 1753. Polypodium bulbiferum L. 1806. Cystopteris bulbifera (L.) Bernh. 1900. Filix bulbifera (L.) Underw.	Bulblet fern, Bulblet bladder-fern, Bulblet fragile-fern, Berry fern Cystoptère bulbifère, Cystoptèride bulbifère	Native, C:8, Secure, DEC-D[1] Perennial, Herb-fern			NYFA-5 USDA-NN NPT Can ARS ITIS FNA  Images, wsp
Cystopteris fragilis  (L.) Bernh. 1753. Polypodium fragile L. 1806. Cystopteris fragilis (L.) Bernh. 1848. Cystopteris dickieana  R.Sim 1900. Filix fragilis (L.) Underw.	Brittle bladder-fern, Fragile fern	Native, C:7, Likely secure, DEC-D[1] Perennial, Herb-fern			NYFA-4 USDA-NN NPT  Images, wsp
Cystopteris tenuis  (Michx.) Desv.   Cystopteris protrusa ×   "Cystopteris hemifragilis"† 1803. Nephrodium tenue Michx. 1827. Cystopteris tenuis (Michx.) Desv. 1889. C. fragilis var. mackayi G.Lawson	Mackay's bladder-fern, Mackay's fragile fern, Upland brittle bladder-fern Cystoptère ténue	Native, C:7, Likely secure, DEC-D[1] Perennial, Herb-fern			NYFA-4 USDA-NN NPT  Images, wsp
Cystopteris protrusa  (Weath.) Blasdell 1935. C. fragilis var. protrusa Weath. 1963. Cystopteris protrusa Blasdell	Lowland bladder-fern, Lowland brittle fern, Lowland fragile fern, Southern bladder-fern	Native, C:10, Endangered, DEC-A[2] Perennial, Herb-fern			NYFA-1 USDA-NN NPT  FNA  Images, wsp
Cystopteris (excluded taxa)	Bladder-fern	N.Y. Status	Images	Distribution	NY NPT
Cystopteris bulbifera × fragilis   Cystopteris bulbifera ×   Cystopteris fragilis	Hybrid of  bulblet bladder-fern &  fragile fern	Native, N.Y. excluded Perennial, Herb-fern			NYFA-XCLD Images, wsp
1 2 3 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered.

Gymnocarpium

Polypodiales — Aspleniaceae — Cystopteridoideae — Gymnocarpium
Gymnocarpium	Oak-fern	N.Y. Status	Images	Distribution	NY NPT
Gymnocarpium dryopteris (L.) Newman 1753. Polypodium dryopteris L. 1851. G. dryopteris (L.) Newman 1852. Phegopteris dryopteris (L.) Fée 1905. Dryopteris linnaeana C. Chr. 1909. Dryopteris dryopteris (L.) Christ 1917. Thelypteris dryopteris (L.) Sloss. 1966. Carpogymnia dryopteris Á.& D.Löve	Northern oak fern, Common oak fern, Western oak fern Gymnocarpe  fougère-du-chêne	Native, C:8, Likely secure, DEC-D[1]			NYFA-4-5 USDA-NN NPT Can ARS ITIS FNA  Images, wsp
Gymnocarpium × brittonianum (Sarvela) Pryer & Haufler   Gymnocarpium disjunctum ×   Gymnocarpium dryopteris 1980. G. dryopteris nssp. brittonianum 1993. G. × brittonianum Pryer & Haufler	Britton's fern, Britton's oak-fern Gymnocarpe de Britton	Native, Unranked			NYFA-U USDA-NN Can ITIS FNA  Images, wsp
Gymnocarpium (excluded)	Oak-fern	N.Y. Status	Images	Distribution	NY NPT
Gymnocarpium disjunctum (Rupr.) Ching 1845. Polypodium dryopteris var. disjunctum 1965. G. disjunctum (Rupr.) Ching	Pacific oakfern, Western oak fern Gymnocarpe disjoint, Dryoptère disjointe, Dryoptèride du chêne	N. America native, N.Y. excluded			NYFA-XCLD USDA-NN Can ITIS FNA Trop. Images, wsp
Gymnocarpium robertianum (Hoffm.) Newman 1795. Polypodium robertianum Hoffm. 1850. Phegopteris robertianum (Hoffm.) Fée 1851. G. robertianum (Hoffm.) Newman 1905. Dryopteris robertiana (Hoffm.) C.Chr. 1917. Thelypteris robertiana (Hoffm.) Sloss. ex Rydb. 1962. G. dryopteris var. pumilum	Scented oakfern, Limestone fern, Robert's oak fern, Northern oak fern Gymnocarpe de Robert	N. America native, N.Y. excluded			NYFA-XCLD USDA-NN Can ITIS FNA Trop. Images, wsp
↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.

Subfamily Asplenioideae

The Asplenioideae Link (1841) (spleenworts)...

Asplenium

Asplenium platyneuron ebony spleenwort

Polypodiales — Aspleniaceae — Asplenioideae — Asplenium
Asplenium	Spleenwort	N.Y. Status	Images	Distribution	NY NPT
Asplenium platyneuron var. platyneuron	Ebony spleenwort	Native, C:5, Secure, DEC-D[1]			NYFA-5 USDA-NN NPT  Images, wsp
Asplenium rhizophyllum  L. 1753. Asplenium rhizophyllum L. 1833. Camptosorus rhizophyllus (L.) Link	Walking fern, Walking-fern spleenwort, American walking fern Doradille ambulante, Camptosore à  feuilles radicantes	Native, C:10, Likely secure, DEC-D[1]			NYFA-4-5 USDA-NN NPT Can ARS ITIS Images, wsp
Asplenium trichomanes  L. ssp. trichomanes	Maidenhair spleenwort	Native, C:9, Likely secure, DEC-D[1]			NYFA-4 USDA-NN NPT  Images, wsp
Asplenium trichomanes  L. ssp. quadrivalens D.E.Mey.	Maidenhair spleenwort	Native, C:9, Likely secure, DEC-D[1]			NYFA-4 USDA-NN NPT  Images, wsp
Asplenium trichomanes  L. nssp. lusaticum (D.E.Mey.) Lawalrèe   A. t. ssp. trichomanes ×   A. t. ssp. quadrivalens 1958. Asplenium × lusaticum D.E.Mey. 1978. A. trichomanes nssp lusaticum	Hybrid  Maidenhair spleenwort	Native, Unranked, No NY reports			NPT  Trop. Images, wsp
Asplenium ruta-muraria  L. 1753. Asplenium ruta muraria L. 1928. Asplenium cryptolepis Fernald 1928. A. cryptolepis var. ohionis Fernald 1935. A. ruta-muraria var. cryptolepis C. Chr. ex Massey 1939. A. ruta-muraria ssp. cryptolepis R.T. Clausen & Wahl	Wallrue, Wall-rue spleenwort Doradille des murailles, Mauerraute	Native, C:10, Likely secure, DEC-D[1]			NYFA-4 USDA-NN NPT Can ARS ITIS FNA Trop. Images, wsp
Asplenium montanum	Mountain spleenwort	Native, C:10, Threatened, DEC-B[2]			NYFA-2-3 USDA-N0 NPT  Images, wsp
Asplenium scolopendrium  L. var. americanum (Fernald) Kartesz & Gandhi 1793. Scolopendrium vulgare Small 1935. Phyllitis scolopendrium var. americana Fernald 1954. Phyllitis fernaldiana Á. Löve 1973. Phyllitis japonica ssp. americana 1991. A. scolopendrium var. americanum	American  hart's-tongue fern, Hart's-tongue fern, Hart's tonguefern	Native, C:9, Threatened, DEC-B[2]			NYFA-2 USDA-N ARS  FNA Trop. Images, wsp
Asplenium viride  Huds. 1753. Asplenium ramosum L. 1753. A. trichomanes-ramosum L. 1762. Asplenium viride Huds.	Green spleenwort, Bright-green spleenwort	Native, C:10, Endangered, DEC-A[3]			NYFA-1 USDA-NN NPT  Trop. Images, wsp
Asplenium bradleyi  D.C.Eaton   Asplenium montanum ×   Asplenium platyneuron 1873. Asplenium bradleyi D.C.Eaton 1925. Asplenium stotleri Wherry 1931. Chamaefilix bradleyi Farw.	Bradley's spleenwort	Native, C:10, No recent reports, Endangered, DEC-A[3]			NYFA-Z USDA-N0 ITIS FNA  Images, wsp
Asplenium × ebenoides  R.R.Scott   Asplenium rhizophyllum ×   Asplenium platyneuron 1865. Asplenium ebenoides R.R.Scott 1937. × Asplenosorus ebenoides Wherry	Scott's spleenwort, Dragon tail fern, Walking spleenwort	Native, No recent reports			NYFA-Z USDA-N0 NPT  Images, wsp
Asplenium (excluded taxa)	Spleenwort	N.Y. Status	Images	Distribution	NY NPT
Asplenium scolopendrium  L. var. scolopendrium 1753. Asplenium scolopendrium L. 1844. Phyllitis scolopendrium Newman 1881. Scolopendrium    scolopendrium H.Karst.	Hart's-tongue fern	Introduced from  Eurasia,  northern Africa, N.Y. excluded, Cultivated			NYFA-XCLD ARS  Trop. Images, wsp
Asplenium ×‌ clermontae  Syme   Asplenium ruta-muraria ×   Asplenium trichomanes 1886. Asplenium clermontae Syme	Hybrid  Wallrue &  Maidenhair spleenwort	Native, Unranked, No NY reports			NYFA-D USDA-N0 NPT  Images, wsp
1 2 3 4 5 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. 1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (b) Threatened. 1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered.

Subfamily Thelypteridoideae

The Thelypteridoideae C.F.Reed (1968) (marsh ferns)

Phegopteris

Polypodiales — Aspleniaceae — Thelypteridoideae — Phegopteris
Phegopteris	Beech-fern	N.Y. Status	Images	Distribution	NY NPT
Phegopteris connectilis (Michx.) Watt 1753. Polypodium phegopteris L. 1803. Polypodium connectile Michx. 1826. Lastrea phegopteris (L.) Bory 1852. Phegopteris polypodioides Fée 1866. Phegopteris connectilis (Michx.) Watt 1905. Dryopteris phegopteris (L.) C.Chr. 1917. Thelypteris phegopteris (L.) Sloss.	Northern beech fern, Narrow beech fern, Long beech fern Phégoptère à     segments joint, Fougère du hêtre	Native, C:7, Secure, DEC-D[1] FACU Perennial, Fern			NYFA-5 USDA-NN NPT Can ARS ITIS FNA Trop. Images, wsp
Phegopteris hexagonoptera (Michx.) Fée 1803. Polypodium hexagonopterum Michx. 1852. Phegopteris hexagonoptera (Michx.) Fée 1905. Dryopteris hexagonoptera (Michx.) C.Chr. 1919. Thelypteris hexagonoptera (Michx.) Weath.	Broad beech fern, Broad beechfern, Southern beech fern Phégoptère à hexagones, Phégoptéride à hexagones, Thélyptère à hexagones	Native, Likely secure, DEC-D[1]			NYFA-4-5 USDA-NN NPT Can ARS  FNA Trop. IPN LBJ Images, wsp
1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.

Thelypteris

Thelypteris palustris marsh fern As with the hay-scented fern (Dennstaedtia punctilobula) and bracken fern (Pteridium aquilinum), listed above, the New York fern (Thelypteris noveboracensis) is considered to be an "interfering" plant in New York forests. New York ferns produce dense stands that prevent the establishment of tree seedlings.[1] Despite being considered weeds (or conceivably because they are), New York ferns, as well as the other two Thelypteris species, are listed as "exploitably vulnerable" by the New York State Department of Environmental Conservation.[2] ↑ Heather M. Engelman & Ralph D. Nyland (2006). "Interference to Hardwood Regeneration in Northeastern North America: Assessing and Countering Ferns in Northern Hardwood Forests." Northern Journal of Applied Forestry (Society of American Foresters) 23(3):166–175. ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.

Polypodiales — Aspleniaceae — Thelypteridoideae — Thelypteris
Thelypteris	Maiden-fern	N.Y. Status	Images	Distribution	NY NPT
Thelypteris palustris  Schott var. pubescens (G.Lawson) Fernald 1836. Dryopteris thelypteris var. pubescens 1864. Lastrea thelypteris var. pubescens 1864. T. confluens var. pubescens 1910. Dryopteris tremula Christ 1929. T. palustris var. haleana 1929. T. palustris var. pubescens	Marsh fern, Eastern marsh fern, Northern marsh fern Thélyptère des marais     d'Amérique	Native, C:4, Secure, DEC-D[1] FACW Perennial, Herb-fern			NYFA-5 USDA-NN NPT Can ARS ITIS FNA Trop. Images, wsp
Thelypteris noveboracensis (L.) Nieuwl. 1753. Polypodium noveboracense L. 1803. Nephrodium thelypterioides Michx. 1827. Nephrodium novaeboracense Desv. 1848. Dryopteris noveboracensis 1910. Thelypteris noveboracensis 1963. Parathelypteris noveboracensis Ching 1972. Thelypteris thelypterioides Holub	New York fern Fougère de New York, Thélyptéride de New York	Native, C:4, Secure, DEC-D[1] FAC Perennial, Herb-fern			NYFA-5 USDA-NN NPT  ARS  FNA  Images, wsp
Thelypteris simulata (Davenp.) Nieuwl. 1894. Aspidium simulatum Davenp. 1894. Dryopteris simulata Davenp. 1904. Filix simulata (Davenp.) Farw. 1910. T. simulata (Davenp.) Nieuwl. 1976. Parathelypteris simulata Holttum	Massachusetts fern, Bog fern Thélyptère simulatrice	Native, C:8, Vulnerable, DEC-D[1] FACW Perennial, Herb-fern			NYFA-3-4 USDA-N NPT  FNA Trop. Images, wsp
Thelypteris (excluded taxa)	Maiden-fern	N.Y. Status	Images	Distribution	NY NPT
Thelypteris palustris  Schott var. palustris 1834. Thelypteris palustris Schott	Eurasian marsh fern Thélyptère des marais	Introduced from  Eurasia, N.Y. excluded			NYFA-XCLD USDA- Images, wsp
1 2 3 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.

Subfamily Woodsioideae

The Woodsioideae Fomin (1911) (cliff ferns)...

Woodsia

Polypodiales — Aspleniaceae — Woodsioideae — Woodsia
Woodsia	Cliff-fern	N.Y. Status	Images	Distribution	NY NPT
Woodsia ilvensis  (L.) R.Br.     Acrostichum ilvense L.	Rusty cliff fern, Rusty woodsia Woodsie de l'île d'Elbe	Native, Secure, DEC-D[1]			NYFA-5 USDA-NN NPT  Images, wsp
Woodsia obtusa  (Spreng.) Torrey ssp. obtusa	Bluntlobe cliff fern, Blunt-lobe woodsia	Native, Secure, DEC-D[1]			NYFA-5 USDA-NN NPT  Images, wsp
Woodsia alpina  (Bolt.) S.Gray     Acrostichum alpinum     W. alpina var. bellii     W. glabella var. bellii	Alpine cliff fern, Alpine woodsia, Northern woodsia Woodsie alpine	Native, Endangered, DEC-A[2]			NYFA-1 USDA-NN NPT  ITIS Images, wsp
Woodsia glabella R.Br. ex Richardson (1823)     W. alpina var. glabella     W. hyperborea var. glabella	Smooth cliff fern, Smooth woodsia Woodsie glabre	Native, Endangered, DEC-A[2]			NYFA-1 USDA-NN glabella/ NPT  FNA  Images, wsp
Woodsia oregana  D.C.Eaton (1865) ssp. cathcartiana 1908. W. cathcartiana B.L.Rob.     W. oregana var. cathcartiana	Oregon cliff fern, Oregon woodsia, (tetraploid)	Native, Likely extirpated			NYFA-Z USDA-NN NPT  ITIS Images, wsp
1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. 1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered.

Subfamily Blechnoideae

The Blechnoideae ...

Onoclea

The sensitive fern, so called due to its sensitivity to frost, is very common in areas that are wet for part of the year.

Polypodiales — Aspleniaceae — Blechnoideae — Onoclea
Onoclea	Sensitive fern	N.Y. Status	Images	Distribution	NY NPT
Onoclea sensibilis  L. 1753. Onoclea sensibilis L.	Sensitive fern, Bead fern, Polypod brake, Meadow brake Onoclée sensible	Native, C:2, Secure FACW Perennial, Fern			NYFA-5 USDA-NN NPT  ARS ITIS FNA  BNA Images, wsp

Matteuccia

With the transfer of Onoclea (above) to Blechnoideae, Christenhusz & Chase (2014) recommended that Matteuccia be moved back to Onoclea.

Polypodiales — Aspleniaceae — Blechnoideae — Matteuccia
Matteuccia	Ostrich fern	N.Y. Status	Images	Distribution	NY NPT
Matteuccia struthiopteris  (L.) Tod. var. pensylvanica (Willd.) C.V.Morton 1810. Struthiopteris pensylvanica Willd. 1819. Onoclea pensylvanica Sm. 1945. Pteretis pensylvanica Fernald 1950. Matteuccia pensylvanica Raymond 1950. Matteuccia struthiopteris var. pensylvanica C.V.Morton 1961. Onoclea struthiopteris var. pensylvanica B.Boivin	Ostrich fern, Shuttlecock fern, Fiddlehead fern Fougère à l'autruche	Native, C:5, Likely secure, DEC-D[1] FACW-FAC Perennial, Herb-fern			NYFA-4 USDA-NN NPT  ARS ITIS FNA  BNA Images, wsp
Matteuccia (excluded taxa)	Ostrich fern	N.Y. Status	Images	Distribution	NY NPT
Matteuccia struthiopteris  (L.) Tod. var. struthiopteris 1753. Osmunda struthiopteris L. 1866. Matteuccia struthiopteris (L.) Tod.	Ostrich fern, Fougère à l'autruche	Introduced from  Eurasia, N.Y. excluded			NYFA-XCLD USDA-NN NPT  ARS ITIS FNA  BNA Images, wsp
↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.

Anchistea

Polypodiales — Aspleniaceae — Blechnoideae — Anchistea
Anchistea	Chainfern	N.Y. Status	Images	Distribution	NY NPT
Anchistea virginica  (L.) C. Presl 1771. Blechnum virginicum L. 1793. Woodwardia virginica (L.) Sm. 1851. Anchistea virginica (L.) C. Presl	Virginia chainfern	Native, C:9, Secure, DEC-D[1] OBL Perennial, Herb-fern			NYFA-5 USDA-N NPT  Images, wsp
↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.

Lorinseria

Polypodiales — Aspleniaceae — Blechnoideae — Lorinseria
Lorinseria	Chainfern	N.Y. Status	Images	Distribution	NY NPT
Lorinseria areolata  (L.) C. Presl 1753. Acrostichum areolatum L. 1851. Lorinseria areolata (L.) C. Presl 1857. Woodwardia areolata (L.) T. Moore	Netted chainfern	Native, C:10, Vulnerable, DEC-D[1] OBL-FACW Perennial, Herb-fern			NYFA-3 USDA-N Images, wsp
↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.

Subfamily Athyrioideae

The Athyrioideae (lady-ferns)...

Deparia

Deparia acrostichoides silvery spleenwort

Polypodiales — Aspleniaceae — Athyrioideae — Deparia
Deparia	False-spleenwort	N.Y. Status	Images	Distribution	NY NPT
Deparia acrostichoides  (Sw.) M.Kato     Athyrium thelypterioides     Diplazium acrostichoides	Silvery spleenwort, Silver false spleenwort, Silvery glade fern	Native, C:8, Secure, DEC-D[1]			NYFA-5 USDA-NN NPT Can Images, wsp
↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.

Athyrium

Polypodiales — Aspleniaceae — Athyrioideae — Athyrium
Athyrium Roth	Lady-fern	N.Y. Status	Images	Distribution	NY NPT
Athyrium angustum  (Willd.) C. Presl 1810. Aspidium angustum Willd. 1825. Athyrium angustum (Willd.) C.Presl 1864. At. filix-femina var. angustum 1949. At. filix-femina ssp. angustum	Northern lady fern, Common lady fern, Narrow lady fern, Subarctic lady fern	Native, C:5, Secure, DEC-D[1]			NYFA-5 USDA-NN NPT Can ARS ITIS FNA Trop. NSE  IPN  Images, wsp
Athyrium asplenioides (Michx.) A.A.Eaton 1803. Nephrodium asplenioides Michx. 1817. At. asplenioides (Michx.) A.A.Eaton. 1923. At. filix-femina var. asplenioides 1962. At. filix-femina ssp. asplenioides	Southern lady fern, Lowland lady fern	Native, C:7, Vulnerable, DEC-D[1]			NYFA-3 USDA-N0 NPT  ITIS FNA Trop. NSE  Images, wsp
Athyrium (excluded taxa)	Lady-fern	N.Y. Status	Images	Distribution	NY NPT
Athyrium filix-femina (L.) Roth ex Mert. = Athyrium angustum × Athyrium asplenioides	Lady fern, Hybrid of  Northern lady fern &  Southern lady fern	Native, N.Y. excluded			NYFA-XCLD BNA Images, wsp
Athyrium nipponicum (Mett.) Hance 1865. Asplenium nipponicum Mett. 1873. Athyrium nipponicum (Mett.) Hance 2011. Anisocampium niponicum Yea C.Liu, W.L.Chiou & M.Kato (Mett.) [2]	Japanese painted lady fern, Japanese painted fern	Introduced from  eastern Asia, No specimens			NYFA-0 NPT  ARS  Trop. Images, wsp
1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ↑ [ Liu & al. (2011) "Molecular phylogeny and taxonomy of the fern genus Anisocampium (Athyriaceae)". TAXON 60 (3), June 2011: 824–830.]

Homalosorus

Homalosorus pycnocarpos narrow-leaved glade fern

Polypodiales — Aspleniaceae — Athyrioideae — Homalosorus
Homalosorus Sw.	Twin-sorus fern	N.Y. Status	Images	Distribution	NY NPT
Homalosorus pycnocarpos Serm. 1803. Asplenium angustifolium Michx. nom. illeg., hom. non Jacq. 1786 1804. Asplenium pycnocarpon Spreng. 1906. Athyrium pycnocarpon Tidestr. 1938. Diplazium pycnocarpon M.Broun 1977. Homalosorus pycnocarpos Serm.	Glade fern, Narrow-leaved glade fern, Narrow-leaved-spleenwort Athyrie à sores denses, Diplazie à sores denses	Native, C:9, Likely secure, DEC-D[1] FAC Perennial, Fern			NYFA-4 USDA-NN NPT Can ITIS FNA Trop. Images, wsp
↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.

Subfamily Dryopteridoideae

The Dryopteridoideae Link (1841) (woodferns)...

Cyrtomium

Polypodiales — Polypodiaceae — Dryopteridoideae — Cyrtomium
Cyrtomium	Netvein-hollyfern	N.Y. Status	Images	Distribution	NY NPT
Cyrtomium falcatum  (L.f.) C.Presl 1782. Polypodium falcatum L.f. 1783. Polypodium japonicum Houtt. 1801. Aspidium falcatum (L.f.) Sw. 1836. Cyrtomium falcatum (L.f.) C. Presl 1891. Dryopteris falcata (L.f.) Kuntze 1899. Polystichum falcatum (L.f.) Diels 1947. Phanerophlebia falcata (L.f.) Copel.	Japanese holly fern, Japanese netvein hollyfern	Introduced from  Asia, N.Y. excluded			NYFA-XCLD USDA-X ARS  Trop. Images, wsp

Dryopteris

Most Dryopteris (wood fern) species are characterized by sori located near the mid leaf vein. Dryopteris hybrids are very common.

Dryopteris diploid species

These species have 2n:82 chromosmes (twice the Dryopteris base chromosome number of n:41). D. intermedia and D. Goldiana have been shown to be part of a North American "reticulate complex", in which, along with three other Dryopteris species, they are parents to most of the allopolyploid Dryopteris species shown below.[1][2] ↑ Emily B. Sessa, Elizabeth A. Zimmer and Thomas J. Givnish (2012). "Unraveling reticulate evolution in North American Dryopteris (Dryopteridaceae)." BioMed Central Evolutionary Biology 2012, 12:104 doi:10.1186/1471-2148-12-104 ↑ Emily B. Sessa, Elizabeth A. Zimmer and Thomas J. Givnish (2012). "Reticulate evolution on a global scale: A nuclear phylogeny for New World Dryopteris (Dryopteridaceae)." Molecular Phylogenetics and Evolution Volume 64, Issue 3, September 2012, Pages 563–581.

Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris
Dryopteris (diploid)	Wood fern	N.Y. Status	Images	Distribution	NY NPT
Dryopteris marginalis  (L.) A.Gray 1753. Polypodium marginale L. 1848. Dryopteris marginalis A.Gray	Marginal wood fern	Native, Secure, DEC-D[1] Perennial, Herb-fern, Anemochore			NYFA-5 USDA-NN NPT  FNA  Images, wsp
Dryopteris intermedia  (Muhl. ex Willd.) A. Gray 1810. Aspidium intermedium Muhl. ex Willd. 1920. Thelypteris intermedia (Muhl. ex Willd.) House(?) 1825. Nephrodium intermedium (Muhl. ex Willd.) C. Presl 1848. Dryopteris intermedia (Muhl. ex Willd.) A. Gray 1893. D. spinulosa var. intermedia 1950. D. austriaca var. intermedia	Evergreen woodfern, Fancy wood fern, Common wood fern	Native, Secure, DEC-D[1] Perennial, Herb-fern, Anemochore			NYFA-5 USDA-NN NPT  FNA  Images, wsp
Dryopteris goldiana  (Hook. ex Goldie) A.Gray 1822. Aspidium goldieanum Hook. ex Goldie 1848. Dryopteris goldieana A.Gray 1910. Thelypteris goldieana Nieuwl. 1916. Filix goldiana Farw. 1931. Filix-mas goldiana Farw.	Goldie's woodfern, Goldie's shield fern, Giant woodfern Dryoptère de Goldie, Dryoptéride de Goldie, Dryoptéris géant	Native, Likely secure, DEC-D[1] Perennial, Herb-fern, Anemochore			NYFA-4 USDA-NN NPT Can ARS ITIS FNA Trop. NSE  IPN  Images, wsp
Dryopteris fragrans  (L.) Schott	Fragrant cliff wood-fern, Fragrant woodfern	Native, Endangered, DEC-A[2] Perennial, Herb-fern, Anemochore			NYFA-1 USDA-NN NPT  FNA  Images, wsp
1 2 3 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered.

Dryopteris allopolyploid species

These are fertile species of hybrid origin. Most are allotetraploid, derived from diploid Dryopteris parents, including two of the New York diploid species, D. intermedia and D. goldiana, shown above. The other diploid parents, D. expansa and D. ludoviciana, are North American, but not New York natives. A fifth, likely extinct, diploid parent "D. semicristata" has been hypothesized to form the well-supported model used here. The species Dryopteris clintoniana is allohexaploid, resulting from the diploid D. goldiana (above) and the allotetraploid D. Cristata, below. The parentage of the introduced non-NY-native, Dryopteris filix-mas has not been resolved, but it is probably not part of the same North American "reticulate complex" as the rest.[1] ↑ Emily B. Sessa, Elizabeth A. Zimmer and Thomas J. Givnish (2012). "Unraveling reticulate evolution in North American Dryopteris (Dryopteridaceae)." BioMed Central Evolutionary Biology 2012, 12:104 doi:10.1186/1471-2148-12-104

Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris
Dryopteris (allopolyploid)	Woodfern	N.Y. Status	Images	Distribution	NY NPT
Dryopteris cristata  (L.) A.Gray   ♀ D. "semicristata" (SS) ×   ♂ D. ludoviciana (LL) → SSLL 1753. Polypodium cristatum L. 1848. Dryopteris cristata A.Gray	Crested shield-fern, Crested woodfern	Native, Secure, DEC-D[1]			NYFA-5 USDA-NN NPT  FNA  Images, wsp
Dryopteris carthusiana (Vill.) H.P.Fuchs   ♀ D. "semicristata" (SS) ×   ♂ D. intermedia (II) → SSII 1777. Polypodium spinulosum O.F.Muell. 1786. Polypodium carthusianum Vill. 1867. Dryopteris spinulosa Watt 1959. Dryopteris carthusiana H.P.Fuchs	Spinulose shield fern, Spinulose wood fern, Toothed wood fern, Narrow buckler fern Dryoptère spinuleuse	Native, Secure, DEC-D[1]			NYFA-5 USDA-N NPT Can ARS  FNA  Images, wsp
Dryopteris clintoniana (D.C.Eaton) Dowell   ♀ D. cristata (SSLL) ×   ♂ D. goldiana (GG) → SSLLGG 1867. Aspidium cristatum var. clintonianum D.C.Eaton 1893. Dryopteris cristata var. clintoniana Underw. 1906. Dryopteris clintoniana Dowell 1920. Thelypteris clintoniana House	Clinton's wood fern Dryoptère de clinton	Native, Secure, DEC-D[1]			NYFA-5 USDA-NN NPT  FNA Trop. Images, wsp
Dryopteris campyloptera (Kunze) Clarkson   ♀ D. intermedia (II) ×   ♂ D. expansa (EE) → IIEE 1848. Aspidium campylopterum Kunze 1915. Dryopteris austriaca Woyn. ex Schinz & Thell. 1915. D. spinulosa var. americana Fernald 1916. D. dilatata var. americana Benedict 1920. Thelypteris dilatata (Hoffm.) House 1930. D. campyloptera Clarkson	Mountain woodfern, Spreading wood fern, Eastern spreading wood fern Dryoptère arquée	Native, Likely secure, DEC-D[1]			NYFA-4 USDA-N NPT Can ARS ITIS FNA Trop. Images, wsp
Dryopteris celsa  (W.Palmer) Knowlt., Palmer & Pollard ex Small   ♀ D. ludoviciana (LL) ×   ♂ D. goldiana (GG) → LLGG 1899. D. goldiana ssp. celsa W.Palmer 1900. Dryopteris celsa Knowlt. &al. 1938. Dryopteris atropalustris Small 1960. Dryopteris wherryi Crane	Log fern	Native, Endangered, DEC-A[2]			NYFA-1 USDA-N ARS ITIS FNA Trop. Images, wsp
Dryopteris (excluded)	Woodfern	N.Y. Status	Images	Distribution	NY NPT
Dryopteris filix-mas  (L.) Schott 1753. Polypodium filix-mas L. 1801. Nephrodium filix-mas Rich. 1834. Dryopteris filix-mas Schott	Male fern, Male wood-fern	N. America native, N.Y. excluded			NYFA-XCLD USDA-NN NPT  ARS  FNA Trop. NSE BNA Images, wsp
1 2 3 4 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered.

Dryopteris named hybrids

Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris subsect. named hybrids
Dryopteris	Woodfern	N.Y. Status	Images	Distribution	NY NPT
Dryopteris × boottii  (Tuck.) Underw.   D. cristata (SSLL) ×   D. intermedia (II) → ILS 1843. Aspidium boottii Tuck. 1893. Dryopteris boottii (Tuck.) Underw. 1910. Thelypteris boottii (Tuck.) Nieuwl.	Boott's wood fern, Glandular swamp fern	Native, Vulnerable			NYFA-3-5 USDA-NN NPT  ITIS Images, wsp
Dryopteris × triploidea  Wherry   D. carthusiana (IISS) ×   D. intermedia (II) → IIS 1930. D. spinulosa var. fructuosa Trudell 1950. D. austriaca var. fructuosa C.V.Morton 1958. D. intermedia var. fructuosa Wherry 1960. Dryopteris × triploidea Wherry	Triploid woodfern	Native, Vulnerable			NYFA-3 USDA-NN ITIS FNA  Images, wsp
Dryopteris × uliginosa (A.Braun ex Dowell) Druce   D. carthusiana (IISS) ×   D. cristata (SSLL) → ILSS	Spinulose crested wood fern	Native, Vulnerable			NYFA-3 USDA-NN Images, wsp
Dryopteris × slossoniae Wherry ex Lellinger   D. cristata (SSLL) ×   D. marginalis (MM) → LMS 1894. D. cristata × marginalis Davenport 1894. Aspidium cristatum × marginale Davenport 1904. Nephrodium slossonae Hahne 1942. Dryopteris × slossonae Wherry	Crested marginal fern, Miss Slosson's wood fern	Native, Threatened			NYFA-1-3 USDA-NN Images, wsp
Dryopteris × pittsfordensis  Slossen   D. carthusiana (IISS) ×   D. marginalis (MM) → IMS 1904. Dryopteris pittsfordensis Sloss. 1904. Nephrodium pittsfordensis Woolson	Pittsford's wood fern	Native, Endangered			NYFA-1 USDA-NN ITIS Trop. IPN  Images, wsp
Dryopteris × burgessii  Boivin   D. clintoniana (SSLLGG) ×   D. marginalis (MM) → GLMS	Burgess' shield fern	Native, No recent reports			NYFA-Z? USDA-NN Images, wsp
Dryopteris × benedictii  (Farw.) Wherry   D. carthusiana (IISS) ×   D. clintoniana (SSLLGG) → GILSS	Benedict's wood fern	Native, No recent reports			NYFA-Z? USDA-NN Images, wsp
Dryopteris × dowellii  (Farw.) Wherry   D. clintoniana (SSLLGG) ×   D. intermedia (II) → GILS	Dowell's wood fern	Native, No recent reports			NYFA-Z USDA-NN Images, wsp
Dryopteris × mickelii   D. clintoniana (SSLLGG) ×   D. goldiana (GG) → GGLS	Wood fern	Native, Likely extirpated			NYFA-Z USDA-N Images, wsp
Dryopteris × neowherryi  W.H.Wagner   D. goldiana (GG) ×   D. marginalis (MM) → GM	Wood fern	Native, No recent reports			NYFA-Z USDA-NN Images, wsp
Dryopteris × separabilis  Small (pro sp.)   D. celsa (LLGG) ×   D. intermedia (II) → GIL	Glandular log fern	Native, No recent reports			NYFA-Z USDA-N0 Images, wsp
Dryopteris × correllii  W.H.Wagner   D. carthusiana (IISS) ×   D. goldiana (GG) → GIS	Correll's wood fern, Poyser's wood fern	Native, No recent reports			NYFA-? USDA-N0 Images, wsp
Dryopteris × leedsii  Wherry   D. celsa (LLGG) ×   D. marginalis (MM) → GLM	Leeds' wood fern	Native, No recent reports			NYFA-Z USDA-N0 NPT  ITIS Images, wsp

Dryopteris unnamed hybrids

Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris subsect. unnamed hybrids
Dryopteris	Woodfern	N.Y. Status	Images	Distribution	NY NPT
Dryopteris filix-mas × marginalis	Marginal male fern	Native, Threatened			NYFA-1-3 Images, wsp
Dryopteris celsa × goldiana   D. celsa (LLGG) ×   D. goldiana (GG) → GGL	Hybrid wood-fern	Native, No recent reports			NYFA-Z Images, wsp
Dryopteris intermedia × marginalis   D. intermedia (II) ×   D. marginalis (MM) → IM	Glandular marginal fern	Native, No recent reports			NYFA-Z Images, wsp
Dryopteris celsa × cristata   D. celsa (LLGG) ×   D. cristata (SSLL) → GLLS	Hybrid wood-fern	Native, No recent reports			NYFA-Z Images, wsp
Dryopteris carthusiana × celsa   D. carthusiana (IISS) ×   D. celsa (LLGG) → IGLS	Spinulose wood fern	Native, No recent reports			NYFA-Z Images, wsp
Dryopteris goldiana × intermedia   D. goldiana (GG) ×   D. intermedia (II) → GI	Wood fern	Native, No recent reports			NYFA-Z Images, wsp
Dryopteris cristata × goldiana   D. cristata (SSLL) ×   D. goldiana (GG) → GLS	Benedict's wood fern	Native, Unranked			NYFA-U Images, wsp

Polystichum

Polystichum acrostichoides Christmas fern Polystichum Roth (1799) goes by the English vernacular names Christmas fern, holy fern, and sword fern.

Polypodiales — Polypodiaceae — Dryopteridoideae — Polystichum
Polystichum	Hollyfern	N.Y. Status	Images	Distribution	NY NPT
Polystichum acrostichoides (Michx.) Schott 1803. Nephrodium acrostichoides Michx. 1834. Polystichum acrostichoides (Michx.) Schott	Christmas fern, Dagger fern Polystic faux-acrostiche	Native, Secure			NYFA-5 USDA-NN NPT  ARS  FNA Trop. BNA Images, wsp
Polystichum braunii  (Spenner) Fée 1825. Aspidium braunii Spenner 1852. Polystichum braunii Fée	Braun's holly-fern	Native, Likely secure			NYFA-4 USDA-NN NPT  Trop. Images, wsp
Polystichum × potteri  Barrington [ P. acrostichoides × P. braunii ] 1986. Polystichum × potteri Barrington	Hybrid Christmas fern   & Braun's holly-fern	Native, Unranked			NYFA-U USDA-NN Trop. Images, wsp
Polystichum lonchitis  (L.) Roth 1753. Polypodium lonchitis L. 1800. Polystichum lonchitis Roth 1801. Aspidium lonchitis Sw. 1924. Polystichum mohrioides var. lemmonii	Northern holly-fern Polystic faux-lonchitis	Native, Endangered			NYFA-Z USDA-NN FNA  Images, wsp
Polystichum × hagenahii  Cody   Polystichum acrostichoides ×   Polystichum lonchitis 1924. P. lonchitis fo. marginale McColl 1966. P. marginale (McColl) Cody 1968. P. × hagenahii Cody	Hagenah's holly fern, Hagenah's polystichum, Hybrid Christmas fern   & northern holly-fern Polystic de Hagenah	Native, Unranked, Questionable[1]			USDA-NN ITIS FNA  Images, wsp
↑ Flora of North America states that the rare hybrid Polystichum × hagenahii has only been found growing with both parents in Ontario. NYFA does not list it, but USDA-PLANTS does show it in NY as well as PA, listing the NY source: Lellinger, D.B. (1985) A field manual of the ferns and fern-allies of the U.S. and Canada, Smithsonian Institution, Washington, DC.