Phlegmariurus
Phlegmariurus is a genus of lycophyte plants in the family Lycopodiaceae.[1] The genus is recognized in the Pteridophyte Phylogeny Group classification of 2016 (PPG I),[2] but not by some other sources, which keep it in a broadly defined Huperzia.
| Phlegmariurus | |
|---|---|
_plant.jpg.webp) | |
| Phlegmariurus phlegmaria growing in the Cambridge University Botanic Garden | |
Scientific classification  | |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Clade: | Lycophytes |
| Class: | Lycopodiopsida |
| Order: | Lycopodiales |
| Family: | Lycopodiaceae |
| Subfamily: | Huperzioideae |
| Genus: | Phlegmariurus (Herter) Holub[1] |
| Species | |
Taxonomy
The genus was first described in 1909 by Wilhelm Herter as the section Phlegmariurus of the genus Lycopodium. The section was elevated to a genus by Josef Ludwig Holub in 1964.[3]
Within the family Lycopodiaceae, Phlegmariurus is placed in the subfamily Huperzioideae. A phylogenetic study in 2016, employing both molecular and morphological data, concluded that either a one-genus or a three-genus division of the subfamily produced monophyletic taxa. The authors preferred the three-genus division, recognizing Huperzia, Phlegmariurus and Phylloglossum. Their preferred hypothesis for the relationships of the three genera was:[4]
| Huperzioideae |
| ||||||||||||
The majority of the species formerly placed in a broadly defined Huperzia belong in Phlegmariurus.[4] However, the genera are difficult to separate morphologically, and others have preferred the one-genus division of the subfamily.[1]
Species
As of January 2023, the Checklist of Ferns and Lycophytes of the World accepted over 300 species:[1]
- Phlegmariurus acerosus (Sw.) B.Øllg.
- Phlegmariurus acifolius (Rolleri) B.Øllg.
- Phlegmariurus acutus (Rolleri) B.Øllg.
- Phlegmariurus affinis (Trevis.) B.Øllg.
- Phlegmariurus afromontanus Pic. Serm.
- Phlegmariurus ambrensis (Rakotondr.) A.R.Field & Bauret
- Phlegmariurus amentaceus (B.Øllg.) B.Øllg.
- Phlegmariurus andinus (Rosenst.) B.Øllg.
- Phlegmariurus aqualupianus (Spring) B.Øllg.
- Phlegmariurus arcuatus (B.Øllg.) B.Øllg.
- Phlegmariurus ascendens (Nessel) B.Øllg.
- Phlegmariurus attenuatus (Spring) B.Øllg.
- Phlegmariurus australis (Willd.) A.R.Field
- Phlegmariurus austroecuadoricus (B.Øllg.) B.Øllg.
- Phlegmariurus austrosinicus (Ching) Li Bing Zhang
- Phlegmariurus axillaris (Roxb.) comb. ined., currently Huperzia axillaris (Roxb.) Rothm.
- Phlegmariurus badinianus (B.Øllg. & P.G.Windisch) B.Øllg.
- Phlegmariurus balansae (Herter) A.R.Field & Bostock
- Phlegmariurus bampsianus (Pic. Serm.) A.R.Field & Bostock
- Phlegmariurus banayanicus (Herter) A.R.Field & Bostock
- Phlegmariurus beitelii (B.Øllg.) B.Øllg.
- Phlegmariurus biformis (Hook.) B.Øllg.
- Phlegmariurus billardierei (Spring) Trevis.
- Phlegmariurus binervius (Herter) B.Øllg.
- Phlegmariurus bolanicus (Rosenst.) A.R.Field & Bostock
- Phlegmariurus brachiatus (Maxon) B.Øllg.
- Phlegmariurus brachystachys (Baker) A.R.Field & Bostock
- Phlegmariurus bradeorum (Christ) B.Øllg.
- Phlegmariurus brevifolius (Hook. & Grev.) B.Øllg.
- Phlegmariurus brongniartii (Spring) B.Øllg.
- Phlegmariurus buesii (Herter) B.Øllg.
- Phlegmariurus callitrichifolius (Mett.) B.Øllg.
- Phlegmariurus campianus (B.Øllg.) B.Øllg.
- Phlegmariurus cancellatus (Spring) Ching
- Phlegmariurus capellae (Herter) B.Øllg.
- Phlegmariurus capillaris (Sodiro) B.Øllg.
- Phlegmariurus carinatus (Poir.) Ching
- Phlegmariurus catacachiensis (Nessel) B.Øllg.
- Phlegmariurus cavifolius (C.Chr.) A.R.Field & Bostock
- Phlegmariurus chamaeleon (Herter) B.Øllg.
- Phlegmariurus changii T.Y.Hsieh
- Phlegmariurus chiricanus (Maxon) B.Øllg.
- Phlegmariurus christii (Silveira) B.Øllg.
- Phlegmariurus ciliatospiculatus B.Øllg.
- Phlegmariurus ciliolatus (B.Øllg.) B.Øllg.
- Phlegmariurus cleefianus (B.Øllg.) B.Øllg.
- Phlegmariurus cocouyensis B.Øllg.
- Phlegmariurus colanensis (B.Øllg.) B.Øllg.
- Phlegmariurus columnaris (B.Øllg.) B.Øllg.
- Phlegmariurus comans (Herter ex Nessel) B.Øllg.
- Phlegmariurus compactus (Hook.) B.Øllg.
- Phlegmariurus copelandianus (R.C.Y.Chou & Bartlett) A.R.Field
- Phlegmariurus coralium (Spring) A.R.Field & Bostock
- Phlegmariurus costaricensis (Herter) comb. ined., currently Huperzia costaricensis (Herter) Holub
- Phlegmariurus crassus (Humb. & Bonpl. ex Willd.) B.Øllg.
- Phlegmariurus creber (Alderw.) A.R.Field & Bostock
- Phlegmariurus crucis-australis (Herter) B.Øllg.
- Phlegmariurus cruentus (Spring) B.Øllg.
- Phlegmariurus cryptomerinus (Maxim.) Satou
- Phlegmariurus cuernavacensis (Underw. & F.E.Lloyd) B.Øllg.
- Phlegmariurus cumingii (Nessel) B.Øllg.
- Phlegmariurus cuneifolius (Hieron.) B.Øllg.
- Phlegmariurus cunninghamioides (Hayata) Ching
- Phlegmariurus curiosus (Herter) A.R.Field & Bostock
- Phlegmariurus curvifolius (Kunze) B.Øllg.
- Phlegmariurus dacrydioides (Baker) A.R.Field & Bostock
- Phlegmariurus dalhousieanus (Spring) A.R.Field & Bostock
- Phlegmariurus darwinianus (Herter ex Nessel) B.Øllg.
- Phlegmariurus delbrueckii (Herter) A.R.Field & Bostock
- Phlegmariurus deminuens (Herter) B.Øllg.
- Phlegmariurus dentatus (Herter) Arana
- Phlegmariurus dianae (Herter) B.Øllg.
- Phlegmariurus dichaeoides (Maxon) B.Øllg.
- Phlegmariurus dichotomus (Jacq.) W.H.Wagner
- Phlegmariurus dielsii (Herter) A.R.Field & Bostock
- Phlegmariurus divergens (Alderw.) A.R.Field & Testo
- Phlegmariurus durus (Pic. Serm.) A.R.Field & Bostock
- Phlegmariurus echinatus (Spring) B.Øllg.
- Phlegmariurus ellenbeckii (Nessel) A.R.Field & Bostock
- Phlegmariurus elmeri (Herter) A.R.Field & Bostock
- Phlegmariurus engleri (Hieron. & Herter) B.Øllg.
- Phlegmariurus eremorum (Rolleri) B.Øllg.
- Phlegmariurus ericifolius (C.Presl) B.Øllg.
- Phlegmariurus erythrocaulos (Fée) B.Øllg.
- Phlegmariurus espinosanus (B.Øllg.) B.Øllg.
- Phlegmariurus eversus (Poir.) B.Øllg.
- Phlegmariurus fargesii (Herter) Ching
- Phlegmariurus filicaulos (Copel.) A.R.Field & Testo
- Phlegmariurus filiformis (Sw.) W.H.Wagner
- Phlegmariurus firmus (Mett.) B.Øllg.
- Phlegmariurus flagellaceus (Kuhn) A.R.Field & Bostock
- Phlegmariurus flexibilis (Fée) B.Øllg.
- Phlegmariurus foliaceus (Maxon) B.Øllg.
- Phlegmariurus foliosus (Copel.) A.R.Field & Bostock
- Phlegmariurus fontinaloides (Spring) B.Øllg.
- Phlegmariurus fordii (Baker) Ching
- Phlegmariurus friburgensis (Herter ex Nessel & Hoehne) B.Øllg.
- Phlegmariurus funiculus (Herter) comb. ined., currently Huperzia funicula (Herter) J.P.Roux
- Phlegmariurus funiformis (Cham. ex Spring) B.Øllg.
- Phlegmariurus gagnepainianus (Herter) A.R.Field & Bostock
- Phlegmariurus galapagensis (O.J.Hamann) B.Øllg.
- Phlegmariurus giganteus (Herter) A.R.Field & Bostock
- Phlegmariurus gnidioides (L. fil.) A.R.Field & Bostock
- Phlegmariurus goebelii (Nessel) A.R.Field & Bostock
- Phlegmariurus gracilis A.Rojas & R.Calderón
- Phlegmariurus guandongensis Ching
- Phlegmariurus gunturensis (Alderw.) A.R.Field & Bostock
- Phlegmariurus haeckelii (Herter) A.R.Field & Testo
- Phlegmariurus haitensis (Herter) comb. ined., currently Huperzia haitensis (Herter) Holub
- Phlegmariurus hamiltonii (Spreng.) Á.Löve & D.Löve
- Phlegmariurus harmsii (Nessel) A.R.Field & Bostock
- Phlegmariurus hartwegianus (Spring) B.Øllg.
- Phlegmariurus hastatus (B.Øllg.) B.Øllg.
- Phlegmariurus hellwigii (Warb.) A.R.Field & Bostock
- Phlegmariurus hemleri (Nessel) B.Øllg.
- Phlegmariurus henryi (Baker) Ching
- Phlegmariurus heterocarpos (Fée) B.Øllg.
- Phlegmariurus heteroclitus (Desv. ex Poir.) B.Øllg.
- Phlegmariurus hexastichus (B.Øllg. & P.G.Windisch) B.Øllg.
- Phlegmariurus hildebrandtii (Herter) A.R.Field & Bauret
- Phlegmariurus hippurideus (Christ) B.Øllg.
- Phlegmariurus hippuris (Desv. ex Poir.) A.R.Field & Testo
- Phlegmariurus hoffmannii (Maxon) B.Øllg.
- Phlegmariurus hohenackeri (Herter) B.Øllg.
- Phlegmariurus holstii (Hieron.) A.R.Field & Bostock
- Phlegmariurus homocarpus (Herter) B.Øllg.
- Phlegmariurus horizontalis (Nessel) A.R.Field & Bostock
- Phlegmariurus huberi (B.Øllg.) B.Øllg.
- Phlegmariurus hugoi B.Øllg.
- Phlegmariurus humbertii (Nessel) A.R.Field & Bostock
- Phlegmariurus humbertii-henrici (Herter) A.R.Field & Bostock
- Phlegmariurus hypogaeus (B.Øllg.) B.Øllg.
- Phlegmariurus hystrix (Herter) B.Øllg.
- Phlegmariurus idroboi B.Øllg.
- Phlegmariurus iminii Kiew
- Phlegmariurus insularis (Carmich.) comb. ined., currently Huperzia insularis (Carmich.) Rothm.
- Phlegmariurus intermedius (Trevis.) B.Øllg.
- Phlegmariurus itambensis (B.Øllg. & P.G.Windisch) B.Øllg.
- Phlegmariurus jaegeri (Herter) A.R.Field & Bostock
- Phlegmariurus josesantae B.Øllg.
- Phlegmariurus kajewskii (Copel.) A.R.Field & Testo
- Phlegmariurus killipii (Herter) B.Øllg.
- Phlegmariurus kuesteri (Nessel) B.Øllg.
- Phlegmariurus lancifolius (Maxon) B.Øllg.
- Phlegmariurus lauterbachii (E.Pritz. ex K.Schum. & Lauterb.) A.R.Field & Bostock
- Phlegmariurus lecomteanus (Nessel) A.R.Field & Bostock
- Phlegmariurus ledermannii (Herter) A.R.Field & Bostock
- Phlegmariurus lehnertii Testo
- Phlegmariurus lignosus (Herter) B.Øllg.
- Phlegmariurus lindenii (Spring) B.Øllg.
- Phlegmariurus linifolius (L.) B.Øllg.
- Phlegmariurus llanganatensis (B.Øllg.) B.Øllg.
- Phlegmariurus lockyeri (D.Jones & B.Gray) A.R.Field & Bostock
- Phlegmariurus loefgrenianus (Silveira) B.Øllg.
- Phlegmariurus longus (Copel.) A.R.Field & Bostock
- Phlegmariurus loxensis (B.Øllg.) B.Øllg.
- Phlegmariurus luteynii B.Øllg.
- Phlegmariurus macbridei (Herter) B.Øllg.
- Phlegmariurus macgregorii (Baker) A.R.Field & Bostock
- Phlegmariurus macrostachys (Hook. ex Spring) N.C.Nair & S.R.Ghosh
- Phlegmariurus magnusianus (Herter) A.R.Field & Testo
- Phlegmariurus mandiocanus (Raddi) B.Øllg.
- Phlegmariurus mannii (Hillebr.) W.H.Wagner
- Phlegmariurus marsupiiformis (D.Jones & B.Gray) A.R.Field & Bostock
- Phlegmariurus martii (Wawra) B.Øllg.
- Phlegmariurus megastachyus (Baker) A.R.Field & Bostock
- Phlegmariurus melanesicus (Brownlie) A.R.Field & Testo
- Phlegmariurus merrillii (Herter) A.R.Field & Bostock
- Phlegmariurus mesoamericanus (B.Øllg.) B.Øllg.
- Phlegmariurus mexicanus (Herter) B.Øllg.
- Phlegmariurus mildbraedii (Herter) A.R.Field & Bostock
- Phlegmariurus mingcheensis Ching
- Phlegmariurus minutifolius (Alderw.) A.R.Field & Bostock
- Phlegmariurus mirabilis (Willd.) A.R.Field & Testo
- Phlegmariurus mollicomus (Spring) B.Øllg.
- Phlegmariurus monticola Kiew
- Phlegmariurus mooreanus (Sander ex Baker) B.Øllg.
- Phlegmariurus multifarius (Alderw.) A.R.Field & Bostock
- Phlegmariurus myrsinites (Lam.) B.Øllg.
- Phlegmariurus myrtifolius (G.Forst.) A.R.Field & Bostock
- Phlegmariurus myrtuosus (Spring) B.Øllg.
- Phlegmariurus neocaledonicus (Nessel) A.R.Field & Bostock
- Phlegmariurus nesselii (Brause ex Nessel) B.Øllg.
- Phlegmariurus niligaricus (Spring) A.R.Field & Bostock
- Phlegmariurus nudus (Christ ex Nessel & Hoehne) B.Øllg.
- Phlegmariurus nummularifolius (Blume) Ching
- Phlegmariurus nutans (Brack.) W.H.Wagner
- Phlegmariurus nyalamensis (Ching & S.K.Wu) H.S.Kung & L.B.Zhang
- Phlegmariurus obovalifolius (Bonap.) A.R.Field & Testo
- Phlegmariurus obtusifolius (P.Beauv.) A.R.Field & Bostock
- Phlegmariurus ocananus (Herter) B.Øllg.
- Phlegmariurus oceanianus (Herter) A.R.Field & Bostock
- Phlegmariurus oellgaardii (A.Rojas) B.Øllg.
- Phlegmariurus oltmannsii (Herter ex Nessel) A.R.Field & Bostock
- Phlegmariurus ophioglossoides (Lam.) A.R.Field & Bostock
- Phlegmariurus orizabae (Underw. & F.E.Lloyd) B.Øllg.
- Phlegmariurus ovatifolius (Ching) W.M.Chu ex H.S.Kung & Li Bing Zhang
- Phlegmariurus pachyphyllus (Kuhn ex Herter) A.R.Field & Testo
- Phlegmariurus pachyskelos B.Øllg.
- Phlegmariurus parksii (Copel.) A.R.Field & Bostock
- Phlegmariurus patentissimus (Alderw.) A.R.Field & Bostock
- Phlegmariurus pecten (Baker) A.R.Field & Bostock
- Phlegmariurus perrierianus (Tardieu) A.R.Field & Bostock
- Phlegmariurus pflanzii (Nessel) B.Øllg.
- Phlegmariurus phlegmaria (L.) Holub
- Phlegmariurus phlegmarioides (Gaudich.) A.R.Field & Bostock
- Phlegmariurus phylicifolius (Desv. ex Poir.) B.Øllg.
- Phlegmariurus phyllanthus (Hook. & Arn.) R.D.Dixit
- Phlegmariurus picardae (Krug) B.Øllg.
- Phlegmariurus pichianus (Tardieu) A.R.Field & Bostock
- Phlegmariurus pinifolius (Trevis.) Kiew
- Phlegmariurus pithyoides (Schltdl. & Cham.) B.Øllg.
- Phlegmariurus pittieri (Christ) B.Øllg.
- Phlegmariurus podocarpensis B.Øllg.
- Phlegmariurus polycarpos (Kunze) B.Øllg.
- Phlegmariurus polycladus (Sodiro) comb. ined., currently Huperzia polyclada (Sodiro) Rolleri & Deferrari
- Phlegmariurus polydactylus (B.Øllg.) B.Øllg.
- Phlegmariurus polylepidetorum (B.Øllg.) B.Øllg.
- Phlegmariurus portoricensis (Underw. & F.E.Lloyd) comb. ined., currently Lycopodium portoricense Underw. & F.E.Lloyd
- Phlegmariurus pringlei (Underw. & F.E.Lloyd) B.Øllg.
- Phlegmariurus proliferus (Blume) A.R.Field & Bostock
- Phlegmariurus pruinosus (Hieron. & Herter) B.Øllg.
- Phlegmariurus pseudovarius (Brownlie) Rouhan & A.R.Field
- Phlegmariurus pulcherrimus (Hook. & Grev.) Á. & D.Löve
- Phlegmariurus pungentifolius (Silveira) B.Øllg.
- Phlegmariurus quadrifariatus (Bory) B.Øllg.
- Phlegmariurus recurvifolius (Rolleri) B.Øllg.
- Phlegmariurus reflexus (Lam.) B.Øllg.
- Phlegmariurus regnellii (Maxon) B.Øllg.
- Phlegmariurus ribourtii (Herter) A.R.Field & Bostock
- Phlegmariurus riobambensis (Nessel) B.Øllg.
- Phlegmariurus robustus (Klotzsch) B.Øllg.
- Phlegmariurus rosenstockianus (Herter) B.Øllg.
- Phlegmariurus rostrifolius (Silveira) B.Øllg.
- Phlegmariurus ruber (Cham. & Schltdl.) B.Øllg.
- Phlegmariurus rubricus (Herter) A.R.Field & Bostock
- Phlegmariurus rufescens (Hook.) B.Øllg.
- Phlegmariurus rupicola (Alderw.) A.R.Field & Bostock
- Phlegmariurus sagasteguianus (B.Øllg.) B.Øllg.
- Phlegmariurus salvinioides (Herter) Ching
- Phlegmariurus samoanus (Herter ex Nessel) A.R.Field & Bostock
- Phlegmariurus sarmentosus (Spring) B.Øllg.
- Phlegmariurus saururus (Lam.) B.Øllg.
- Phlegmariurus scabridus (B.Øllg.) B.Øllg.
- Phlegmariurus schlechteri (E.Pritz.) A.R.Field & Bostock
- Phlegmariurus schlimii (Herter) B.Øllg.
- Phlegmariurus schmidtchenii (Hieron.) B.Øllg.
- Phlegmariurus sellifolius (B.Øllg.) B.Øllg.
- Phlegmariurus sellowianus (Herter) B.Øllg.
- Phlegmariurus serpentiformis (Herter) B.Øllg.
- Phlegmariurus setifolius (Alderw.) A.R.Field & Bostock
- Phlegmariurus shangsiensis C.Y.Yang
- Phlegmariurus shingianus R.H.Jiang & X.C.Zhang
- Phlegmariurus sieberianus (Spring) B.Øllg.
- Phlegmariurus sieboldii (Miq.) Ching
- Phlegmariurus silveirae (Herter ex Nessel & Hoehne) B.Øllg.
- Phlegmariurus silverstonei B.Øllg.
- Phlegmariurus sintenisii (Herter) B.Øllg.
- Phlegmariurus societensis (J.W.Moore) A.R.Field
- Phlegmariurus sooianus Lawalrée
- Phlegmariurus sotae (Rolleri) B.Øllg.
- Phlegmariurus sphagnicola B.Øllg.
- Phlegmariurus squarrosus (G.Forst.) Á. & D.Löve
- Phlegmariurus staudtii (Nessel) A.R.Field & Bostock
- Phlegmariurus stemmermanniae A.C.Medeiros & Wagner
- Phlegmariurus stephani B.Øllg.
- Phlegmariurus strictus (Baker) A.R.Field & Bostock
- Phlegmariurus subfalciformis (Alderw.) A.R.Field & Bostock
- Phlegmariurus subtubulosus (Herter) comb. ined., currently Huperzia subtubulosa (Herter) Holub
- Phlegmariurus subulatus (Desv. ex Poir.) B.Øllg.
- Phlegmariurus tafaensis (Nessel) A.R.Field & C.W.Chen
- Phlegmariurus talamancanus (B.Øllg.) B.Øllg.
- Phlegmariurus talamauanus (Alderw.) A.R.Field & Bostock
- Phlegmariurus talpiphilus (B.Øllg.) B.Øllg.
- Phlegmariurus tardieuae (Herter) A.R.Field & Testo
- Phlegmariurus tauri (Herter) A.R.Field & Testo
- Phlegmariurus taxifolius (Sm.) Á. & D.Löve
- Phlegmariurus tenuicaulis (Underw. & F.E.Lloyd) B.Øllg.
- Phlegmariurus tenuis (Humb. & Bonpl. ex Willd.) B.Øllg.
- Phlegmariurus terrae-guilelmii (Herter) A.R.Field & Bostock
- Phlegmariurus tetragonus (Hook. & Grev.) B.Øllg.
- Phlegmariurus tetrastichoides (A.R.Field & Bostock) A.R.Field & Bostock
- Phlegmariurus tetrastichus (Kunze) A.R.Field & Bostock
- Phlegmariurus tico A.Rojas
- Phlegmariurus toppingii (Herter) A.R.Field & Bostock
- Phlegmariurus tournayanus Lawalrée
- Phlegmariurus transilla (Sodiro ex Baker) B.Øllg.
- Phlegmariurus treitubensis (Silveira) B.Øllg.
- Phlegmariurus trifoliatus (Copel.) A.R.Field & Bostock
- Phlegmariurus trigonus (C.Chr.) A.R.Field & Bostock
- Phlegmariurus tryonorum B.Øllg.
- Phlegmariurus tubulosus (Maxon) B.Øllg.
- Phlegmariurus ulixis (Herter) B.Øllg.
- Phlegmariurus unguiculatus (B.Øllg.) B.Øllg.
- Phlegmariurus urbanii (Herter) B.Øllg.
- Phlegmariurus vanuatuensis A.R.Field
- Phlegmariurus varius (R.Br.) A.R.Field & Bostock
- Phlegmariurus venezuelanicus (Herter) B.Øllg.
- Phlegmariurus vernicosus (Hook. & Grev.) Á. & D.Löve
- Phlegmariurus villonacensis (B.Øllg.) B.Øllg.
- Phlegmariurus warneckei (Herter ex Nessel) A.R.Field & Testo
- Phlegmariurus watsonianus (Maxon) B.Øllg.
- Phlegmariurus weberbaueri (Nessel) B.Øllg.
- Phlegmariurus weddellii (Herter) B.Øllg.
- Phlegmariurus wilsonii (Underw. & F.E.Lloyd) B.Øllg.
- Phlegmariurus xiphophyllus (Baker) A.R.Field & Bostock
- Phlegmariurus yunfengii R.H.Jiang & X.C.Zhang
- Phlegmariurus yunnanensis Ching
One hybrid is known:[1]
- Phlegmariurus × koolauensis W.H.Wagner
References
- Hassler, Michael (19 January 2023), "Phlegmariurus", World Ferns. Synonymic Checklist and Distribution of Ferns and Lycophytes of the World, 14.7
- PPG I (2016), "A community-derived classification for extant lycophytes and ferns", Journal of Systematics and Evolution, 54 (6): 563–603, doi:10.1111/jse.12229, S2CID 39980610
- "Phlegmariurus (Herter) Holub", The International Plant Names Index, retrieved 2019-10-14
- Field, Ashley R.; Testo, Weston; Bostock, Peter D.; Holtum, Joseph A. M. & Waycott, Michelle (2016), "Molecular phylogenetics and the morphology of the Lycopodiaceae subfamily Huperzioideae supports three genera: Huperzia, Phlegmariurus and Phylloglossum", Molecular Phylogenetics and Evolution, 94 (Pt B): 635–657, doi:10.1016/j.ympev.2015.09.024, PMID 26493224
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