List of amphibians of India
This is an index to the amphibians found in India. The amphibians of India show a high level of endemism.[1] This list is based largely on Darrel Frost (2006)[2] and includes common names from older books and journals.[3][4][5]
Order Anura
Family Bufonidae
- Ornate torrent toad, Ansonia ornata = Ghatophryne ornata (Günther, 1876)
- Silent Valley torrent toad, Ansonia ornata = Ghatophryne rubigina (Pillai and Pattabiraman, 1981)
- Bufo beddomii (Günther, 1876)
- Bufo brevirostris (Rao, 1937)
- Bufo burmanus (Andersson, 1939)
- Duttaphrynus cyphosus = Bufo cyphosus (Ye, 1977)
- Himalayan toad, Duttaphrynus himalayanus (Günther, 1864) = Bufo himalayanus (Günther, 1864)
- Bufo hololius (Günther, 1876)
- Xanthophryne koynayensis (Soman, 1963)
- Xanthophryne tigerina Biju, Bocxlaer, Giri, Loader and Bossuyt, 2009
- Ladakh toad, Pseudepidalea latastii Boulenger, 1882 = Bufo latastii
- Common Indian toad, Duttaphrynus melanostictus (Schneider, 1799) = Bufo melanostictus Schneider, 1799
- Duttaphrynus microtympanum (Boulenger, 1882) = Bufo microtympanum (Boulenger, 1882)
- Ingerophrynus macrotis (Boulenger, 1887)
- Bufo parietalis (Boulenger, 1882)
- Ferguson's toad, Bufo scaber (Schneider, 1799)
- Bufo silentvalleyensis (Pillai, 1981)
- Bufo stomaticus (Lütken, 1864)
- Green toad, Bufo viridis
- Bufo stuarti (Smith, 1929)
- Bufoides meghalayanus (Yazdani and Chanda, 1971)
- Pedostibes kempi (Boulenger, 1919)
- Pedostibes tuberculosus (Günther, 1876)
Dicroglossinae
- Common skittering frog, Euphlyctis cyanophlyctis (Schneider, 1799)
- Euphlyctis ghoshi (Chanda, 1991)
- Indian five-fingered frog, Euphlyctis hexadactylus (Lesson, 1834)
- Karavali skittering frog, Euphlyctis karaavali Priti et al., 2016
- Fejervarya andamanensis (Stoliczka, 1870) = Limnonectes andamanensis (Stoliczka, 1870)
- Fejervarya assimilis (Blyth, 1852)
- Fejervarya brama (Lesson, 1834)
- Fejervarya brevipalmata (Peters, 1871)
- Fejervarya cancrivora (Gravenhorst, 1829)
- Fejervarya greenii (Boulenger, 1905)
- Fejervarya keralensis (Dubois, 1981)
- Alpine cricket frog, Fejervarya limnocharis (Gravenhorst, 1829)
- Fejervarya murthii (Pillai, 1979)
- Fejervarya mysorensis (Rao, 1922)
- Fejervarya nicobariensis (Stoliczka, 1870)
- Fejervarya nepalensis (Iskandar, 1998)
- Fejervarya nilagirica (Jerdon, 1854)
- Fejervarya orissaensis (Dutta, 1997)
- Fejervarya parambikulamana (Rao, 1937)
- Fejervarya rufescens (Jerdon, 1854)
- Fejervarya sauriceps (Rao, 1937)
- Fejervarya syhadrensis = Limnonectes syhadrensis (Annandale, 1919)
- Fejervarya teraiensis (Dubois, 1984)
- Jerdon's bullfrog, Hoplobatrachus crassus (Jerdon, 1854)
- Indian bullfrog, Hoplobatrachus tigerinus (Daudin, 1802)
- Limnonectes doriae (Boulenger, 1887)
- Limnonectes hascheanus (Stoliczka, 1870)
- Limnonectes khasianus (Anderson, 1871)
- Limnonectes kuhlii (Tschudi, 1838)
- Limnonectes laticeps (Boulenger, 1882)
- Limnonectes limborgi (Sclater, 1892)
- Limnonectes mawlyndipi (Chanda, 1990)
- Limnonectes mawphlangensis (Pillai and Chanda, 1977)
- Limnonectes shompenorum (Das, 1996)
- Minervarya charlesdarwini (Das, 1998) = Rana charlesdarwini
- Minervarya sahyadris (Dubois, Ohler, and Biju, 2001)
- Nanorana annandalii = Paa annandalii (Boulenger, 1920)
- Nanorana arnoldi = Paa arnoldi (Dubois, 1975)
- Nanorana barmoachensis = Paa barmoachensis (Khan and Tasnim, 1989)
- Nanorana blanfordii = Paa blanfordii (Boulenger, 1882)
- Nanorana conaensis = Paa conaensis (Fei and Huang In Huang and Fei, 1981)
- Nanorana ercepeae = Paa ercepeae (Dubois, 1974)
- Hazara torrent frog, Nanorana hazarensis = Paa hazarensis (Dubois and Khan, 1979)
- Nanorana liebigii = Paa liebigii (Günther, 1860)
- Nanorana minica = Paa minica (Dubois, 1975)
- Nanorana mokokchungensis = Paa mokokchungensis (Das and Chanda, 2000)
- Nanorana polunini = Paa polunini (Smith, 1951)
- Nanorana pleskei (Günther, 1896)
- Karez frog, Nanorana sternosignata = Paa sternosignata (Murray, 1885)
- Murree Hills frog, Nanorana vicina = Paa vicina (Stoliczka, 1872)
- Ombrana sikimensis = Chaparana sikimensis (Jerdon, 1870)
- Burrowing frog, Sphaerotheca breviceps (Schneider, 1799)
- Sphaerotheca dobsoni (Boulenger, 1882)
- Sphaerotheca leucorhynchus (Rao, 1937)
- Marbled sand frog Sphaerotheca rolandae (Dubois, 1983)
Occidozyginae
- Occidozyga borealis (Annandale, 1912)
- Occidozyga lima (Gravenhorst, 1829)
- Occidozyga sumatrana (Peters, 1877)
Family Hylidae
- Hyla annectans (Jerdon, 1870)
Family Megophryidae
- Hasselt's toad, Leptobrachium hasseltii (Tschudi, 1838)
- Leptobrachium smithi (Matsui, Nabhitabhata, and Panha, 1999)
- Scutiger boulengeri (Bedriaga, 1898)
- Scutiger gongshanensis (Yang and Su In Yang, Su, and Li, 1979)
- Nyingchi alpine toad, Scutiger nyingchiensis (Fei, 1977)
- Sikkim alpine toad, Scutiger sikimmensis (Blyth, 1855)
- Xenophrys boettgeri (Boulenger, 1899)
- Xenophrys kempii (Annandale, 1912)
- Xenophrys major (Boulenger, 1908)
- Xenophrys parva (Boulenger, 1893)
- Xenophrys robusta (Boulenger, 1908)
- Xenophrys wuliangshanensis
Family Micrixalidae
- Micrixalus elegans (Rao, 1937)
- Micrixalus fuscus (Boulenger, 1882)
- Micrixalus gadgili Pillai and Pattabiraman, 1990
- Micrixalus kottigeharensis (Rao, 1937)
- Micrixalus narainensis (Rao, 1937)
- Micrixalus nudis Pillai, 1978
- Micrixalus phyllophilus (Jerdon, 1854)
- Micrixalus saxicola (Jerdon, 1854)
- Micrixalus silvaticus (Boulenger, 1882)
- Micrixalus swamianus (Rao, 1937)
- Micrixalus thampii (Pillai, 1981)
Family Microhylidae
- Kalophrynus interlineatus (Blyth, 1855) = Kalophrynus orangensis (Dutta, Ahmed, and Das, 2000)
- Micryletta inornata (Boulenger, 1890)
- Ramanella anamalaiensis (Rao, 1937)
- Ramanella minor (Rao, 1937)
- Ramanella montana (Jerdon, 1854)
- Ramanella mormorata (Rao, 1937)
- Ramanella triangularis (Günther, 1876)
- White-bellied pug-snout frog, Ramanella variegata (Stoliczka, 1872)
- Uperodon globulosus (Günther, 1864)
- Marbled balloon frog, Uperodon systoma (Schneider, 1799)
- Melanobatrachus indicus (Beddome, 1878)
- Kaloula baleata (Müller In Oort and Müller, 1833)
- Painted frog, Kaloula pulchra (Gray, 1831)
- Kaloula assamensis (Das, Sengupta, Ahmed, and Dutta, 2005)
- Sri Lankan bullfrog, Kaloula taprobanica (Parker, 1934)
- Microhyla berdmorei (Blyth, 1856)
- Microhyla chakrapanii (Pillai, 1977)
- Microhyla heymonsi (Vogt, 1911)
- Ornate narrow-mouthed toad, Microhyla ornata (Duméril and Bibron, 1841)
- Microhyla pulchra (Hallowell, 1861)
- Microhyla rubra (Jerdon, 1854)
- Microhyla sholigari (Dutta and Ray, 2000)
- Microhyla laterite Seshadri et al., 2016
- Mysticellus franki Garg & Biju, 2019
Family Nyctibatrachidae
- Nyctibatrachus aliciae (Inger, Shaffer, Koshy, and Bakde, 1984)
- Nyctibatrachus beddomii (Boulenger, 1882)
- Nyctibatrachus deccanensis (Dubois, 1984)
- Nyctibatrachus humayuni (Bhaduri and Kripalani, 1955)
- Nyctibatrachus hussaini (Krishnamurthy, Reddy, and Gururaja, 2001)
- Nyctibatrachus kempholeyensis (Rao, 1937)
- Nyctibatrachus major (Boulenger, 1882)
- Nyctibatrachus minor (Inger, Shaffer, Koshy, and Bakde, 1984)
- Nyctibatrachus periyar Biju et al., 2011
- Nyctibatrachus petraeus (Das and Kunte, 2005)
- Nyctibatrachus sanctipalustris (Rao, 1920)
- Nyctibatrachus sylvaticus (Rao, 1937)
- Nyctibatrachus vasanthi (Ravichandran, 1997)
- Nyctibatrachus minimus Biju et al., 2007.[6]
- Nyctibatrachus sholai Radhakrishnan, Dinesh & Ravichandran, 2007.[7]
- Nyctibatrachus dattatreyaensis Dinesh K P, Radhakrishnan C & Gopalakrishna Bhatta, 2008[8]
Family Petropedetidae
- Indirana beddomii (Günther, 1876)
- Indirana brachytarsus (Günther, 1876)
- Indirana diplosticta (Günther, 1876)
- Indirana gundia (Dubois, 1986)
- Indirana leithii (Boulenger, 1888)
- Indirana leptodactyla (Boulenger, 1882)
- Indirana longicrus (Rao, 1937)
- Indirana phrynoderma (Boulenger, 1882)
- Indirana semipalmata (Boulenger, 1882)
- Indirana tenuilingua (Rao, 1937)
Family Ranidae
- Amolops chakrataensis (Ray, 1992)
- Amolops formosus (Günther, 1876)
- Amolops gerbillus (Annandale, 1912)
- Amolops himalayanus (Boulenger, 1888)
- Amolops jaunsari (Ray, 1992)
- Amolops marmoratus (Blyth, 1855)
- Amolops monticola (Anderson, 1871)
- Amolops viridimaculatus (Jiang, 1983)
- Bicolored frog, Clinotarsus curtipes (Jerdon, 1854)
- Huia chloronota (Günther, 1876)
- Humerana humeralis (Boulenger, 1887)
- Fungoid frog, Hydrophylax malabaricus (Tschudi, 1838)
- Hydrophylax bahuvistara
- Hydrophylax raniceps (Peters, 1871)
- Pterorana khare (Kiyasetuo and Khare, 1986)
- Hylarana tytleri (Theobald, 1868)
- Nasirana alticola (Boulenger, 1882)
- Golden frog, Hylarana aurantiaca (Boulenger, 1904)
- Hylarana chitwanensis (Das, 1998)
- Boulenger's Garo hill frog, Hylarana garoensis (Boulenger, 1920 )
- Hylarana leptoglossa (Cope, 1868)
- Hylarana nigrovittata (Blyth, 1856)
- Bronzed frog, Hylarana temporalis (Günther, 1864)
Family Rhacophoridae
- Chiromantis cherrapunjiae = Chirixalus cherrapunjiae (Roonwal and Kripalani, 1966)
- Chiromantis doriae = Chirixalus doriae (Boulenger, 1893)
- Chiromantis dudhwaensis = Chirixalus dudhwaensis (Ray, 1992)
- Chiromantis shyamrupus = Chirixalus shyamrupus (Chanda and Ghosh, 1989)
- Chiromantis simus = Chirixalus simus (Annandale, 1915)
- Chiromantis vittatus = Chirixalus vittatus (Boulenger, 1887)
- Ghatixalus variabilis (Jerdon, 1853)[9]
- Nyctixalus moloch (Annandale, 1912)
- Kurixalus verrucosus = Rhacophorus verrucosus (Boulenger, 1893)
- Aquixalus bisacculus = Rhacophorus bisacculus (Taylor, 1962)
- Aquixalus naso = Polypedates naso (Annandale, 1912)
- Raorchestes andersoni (Ahl, 1927)
- Raorchestes anili (Biju and Bossuyt, 2006)
- Raorchestes luteolus (Kuramoto and Joshy, 2003)
- Raorchestes tuberohumerus (Kuramoto and Joshy, 2003)
- Raorchestes annandalii (Boulenger, 1906)
- Raorchestes beddomii (Günther, 1876)
- Raorchestes bobingeri (Biju and Bossuyt, 2005)
- Raorchestes bombayensis (Annandale, 1919)
- Raorchestes chalazodes (Günther, 1876)
- Raorchestes charius (Rao, 1937)
- Raorchestes dubius (Boulenger, 1882)
- Raorchestes femoralis Round-snout pygmy frog (Günther, 1864)
- Raorchestes flaviventris (Boulenger, 1882)
- Raorchestes garo (Boulenger, 1919)
- Raorchestes glandulosus (Jerdon, 1854)
- Raorchestes graminirupes (Biju & Bossuyt, 2005)
- Raorchestes griet (Bossuyt, 2002)
- Raorchestes jerdonii (Günther, 1876)
- Raorchestes kempiae (Boulenger, 1919)
- Raorchestes luteolus = Raorchestes neelanethrus Gururaja et al., 2007[10]
- Raorchestes microdiscus (Annandale, 1912)
- Raorchestes namdaphaensis (Sarkar and Sanyal, 1985)
- Sharp-snout pygmy tree frog, Raorchestes nasutus
- Raorchestes nerostagona
- Raorchestes ochlandrae Gururaja et al., 2007.[11]
- Raorchestes ponmudi
- Raorchestes sanctisilvaticus (Das and Chanda, 1997)
- Raorchestes shillongensis (Pillai and Chanda, 1973)
- Raorchestes signatus (Boulenger, 1882)
- Raorchestes similipalensis (Dutta, 2003)
- Raorchestes terebrans (Das and Chanda, 1998)
- Raorchestes tinniens (Jerdon, 1854)
- Raorchestes travancoricus (Boulenger, 1891)
- Raorchestes tuberohumerus
- Raorchestes wynaadensis (Jerdon, 1854)
- Polypedates bengalensis Purkayastha et al., 2019[12]
- Polypedates gongshanensis
- Polypedates insularis (Das, 1995)
- Polypedates leucomystax (Gravenhorst, 1829)
- Polypedates maculatus (Gray, 1833) chunam frog
- Polypedates megacephalus (Hallowell, 1861)
- Polypedates pseudocruciger (Das and Ravichandran, 1998)
- Polypedates taeniatus (Boulenger, 1906)
- Rhacophorus appendiculatus (Günther, 1858)
- Rhacophorus bipunctatus (Ahl, 1927)
- Rhacophorus calcadensis (Ahl, 1927)
- Rhacophorus lateralis (Boulenger, 1883)
- Malabar flying frog, Rhacophorus malabaricus (Jerdon, 1870)
- Rhacophorus maximus (Günther, 1858)
- Rhacophorus namdaphaensis (Sarkar and Sanyal, 1985)
- Rhacophorus pseudomalabaricus (Vasudevan and Dutta, 2000)
- Rhacophorus tuberculatus (Anderson, 1871)
- Rhacophorus variabilis (Jerdon, 1854)
- Rhacophorus htunwini (Wilkinson, Thin, Lwin, and Shein, 2005)
- Rhacophorus translineatus (Wu In Sichuan Institute of Biology Herpetology Department, 1977)
- Rhacophorus kio (Ohler and Delorme, 2006)
- Theloderma asperum (Boulenger, 1886)
Family Sooglossidae
- Purple frog, Nasikabatrachus sahyadrensis (Biju and Bossuyt, 2003)
- Bhupathy's purple frog, Nasikabatrachus bupathi (Janani, Vasudevan, Prendini, Dutta, and Aggarwal, 2017)
Order Gymnophiona
The list of Indian caecilians is based on Giri & Gaikwad (2013) (duly amended):[13]
Family Caeciliidae
- Gegeneophis spp.
- Gegeneophis carnosus (Beddome, 1870)
- Amboli caecilian, Gegeneophis danieli Giri, Gower and Wilkinson 2003
- Gegeneophis goaensis Bhatta, Dinesh, Prashanth & Kulkarni, 2007
- Gegeneophis krishni Pillai and Ravichandran, 1999
- Gegeneophis madhavai Bhatta and Srinivasa, 2004[14]
- Gegeneophis mhadeiensis Bhatta, Dinesh, Prashanth & Kulkarni, 2007
- Gegeneophis nadkarnii Bhatta and Prashanth, 2004[15]
- Gegeneophis pareshi Giri, Gower, Gaikwad & Wilkinson, 2011
- Gegeneophis primus Kotharambath, Gower, Oomen & Wilkinson, 2012
- Forest caecilian, Gegeneophis ramaswamii Taylor, 1964
- Gegeneophis seshachari Ravichandran, Gower, and Wilkinson, 2003
- Indotyphlus spp.
- Indotyphlus battersbyi Taylor, 1960
- Maharashtra caecilian, Indotyphlus maharashtraensis Giri, Wilkinson, and Gower, 2003
Family Chikilidae
- Fuller's caecilian, Chikila fulleri (Alcock, 1904) formerly Gegeneophis fulleri[16]
- Chikila alcocki Kamei, Gower, Wilkinson & Biju, 2013[17]
- Chikila darlong Kamei, Gower, Wilkinson & Biju, 2013[17]
- Chikila gaiduwani Kamei, Gower, Wilkinson & Biju, 2013[17]
Family Ichthyophiidae
Ichthyophis spp.[18]
- Ichthyophis alfredii Mathew & Sen, 2009
- Ichthyophis beddomei Peters, 1880
- Ichthyophis bombayensis Taylor, 1960
- Ichthyophis daribokensis Mathew & Sen 2009
- Ichthyophis garoensis Pillai and Ravichandran, 1999
- Ichthyophis davidi Bhatta, Dinesh, Kulkarni, Radhakrishnan 2011
- Ichthyophis husaini Pillai and Ravichandran, 1999
- Ichthyophis kodaguensis Wilkinson, Gower, Govindappa & Venkatachalaiah, 2007
- Ichthyophis khumhzi Kamei, Wilkinson, Gower & Biju, 2009
- Ichthyophis longicephalus Pillai, 1986
- Ichthyophis moustakius Kamei, Wilkinson, Gower & Biju, 2009
- Ichthyophis nokrekensis Mathew & Sen, 2009
- Ichthyophis sendenyu Kamei, Wilkinson, Gower & Biju, 2009
- Ichthyophis sikkimensis Taylor, 1960
- Ichthyophis tricolor Annandale, 1909
- Uraeotyphlus spp.
- Uraeotyphlus gansi Gower, Rajendran, Nussbaum, & Wilkinson, 2008
- Uraeotyphlus interruptus Pillai and Ravichandran, 1999
- Uraeotyphlus malabaricus (Beddome, 1870)
- Uraeotyphlus menoni Annandale, 1913
- Uraeotyphlus narayani Seshachar, 1939
- Uraeotyphlus oommeni Gower & Wilkinson, 2007
- Uraeotyphlus oxyurus (Duméril and Bibron, 1841)
References
- Daniels, R. J. R. (2001). "Endemic fishes of the Western Ghats and the Satpura hypothesis" (PDF). Current Science. 81 (3): 240–244.
- Frost, Darrel R. (17 August 2006). "Amphibian Species of the World: An Online Reference" (Electronic database). 4. American Museum of Natural History. Archived from the original on 24 October 2007. Retrieved 16 November 2007.
- Daniels, R. J. Ranjit (2005). Amphibians of Peninsular India. Universities Press.
- Chanda, Shyamal Kumar; Das, Indraneil & Dubois, Alain (2000). "Catalogue of amphibian types in the collection of the Zoological Survey of India". Hamadryad. 25 (2): 100–128. Archived from the original on 20 July 2011
- Sclater, William Lutley (1892). List of the Batrachia in the Indian Museum.
- Biju, S. D. et al. (2007). "A new nightfrog Nyctibatrachus minimus sp. nov. (Anura: Nyctibatrachidae): The smallest frog from India". Current Science. 93 (6): 854-858.
- Radhakrishnan, C.; Dinesh, K. P. & Ravichandran, M. S. (2007). "A new species of Nyctibatrachus Boulenger (Amphibia: Anura: Nyctibatrachidae) from the Eravikulam National Park, Kerala, India". Zootaxa. 1595: 31–41.
- Dinesh, K. P.; C, Radhakrishnan & Bhatta, Gopalakrishna (2008). "A new species of Nyctibatrachus Boulenger (Amphibia: Anura: Nyctibatrachidae) from the surroundings of Bhadra Wildlife Sanctuary, Western Ghats, India", Zootaxa. 1914: 45–56.
- Biju, S.D.; Roelants, Kim & Bossuyt, Franky (2008). "Phylogenetic position of the montane treefrog Polypedates variabilis Jerdon, 1853 (Anura: Rhacophoridae), and description of a related species". Organisms, Diversity & Evolution. 8: 267–276.
- Gururaja, K. V.; Aravind, N. A.; Ali, S.; Ramachandra, T. V.; Velavan, T. P.; Krishnakumar, V. & Aggarwal, R. K. (2007). "A New Frog Species from the Central Western Ghats of India, and its Phylogenetic Position". Zoological Science. 24: 525-534.
- Gururaja, K. V.; Dinesh, K. P.; Palot, M. J.; Radhakrishnan, C. & Ramachandra, T. V. (2007). "A new species of Raorchestes Gistel (Amphibia: Anura: Rhacophoridae) from southern Western Ghats, India". Zootaxa. 1621: 1-16.
- Purkayastha, Jayaditya; Das, Madhurima; Mondal, Kingshuk; Mitra, Shibajee; Chaudhuri, Anirban & Das, Indraneil (2019). "A new species of Polypedates Tschudi, 1838 (Amphibia: Anura: Rhacophoridae) from West Bengal State, Eastern India". Zootaxa. 4691 (5): 525–540. doi:10.11646/zootaxa.4691.5.6.
- Giri, V. B. & Gaikwad, K. S. (2013). Towards the Conservation of Caecilian Amphibians of the Northern Western Ghats: Establishing the Systematic Foundation. Mumbai: Bombay Natural History Society. p. 53.
- Bhatta, Gopalakrishna & Srinivasa, R. (2004). "A new species of Gegeneophis Peters (Amphibia: Gymnophiona: Caeciliidae) from the surroundings of Mookambika Wildlife Sanctuary, Karnataka, India". Zootaxa. 644: 1–8.
- Bhatta, Gopalakrishna & Prashanth, P. (2004). "Gegeneophis nadkarnii – a caecilian (Amphibia: Gymnophiona: Caeciliidae) from Bondla Wildlife Sanctuary, Western Ghats". Current Science. 87 (3): 10.
- Kamei, R. G.; Mauro, D. S.; Gower, D. J.; Van Bocxlaer, I.; Sherratt, E.; Thomas, A.; Babu, S.; Bossuyt, F.; Wilkinson, M. & Biju, S. D. (22 February 2012). "Discovery of a new family of amphibians from northeast India with ancient links to Africa". Proceedings of the Royal Society B: Biological Sciences. 279 (1737): 2396–2401. doi:10.1098/rspb.2012.0150. ISSN 0962-8452. PMC 3350690. PMID 22357266.
- Kamei, R. G.; Gower, D. J.; Wilkinson, M. & Biju, S. D. (4 June 2013). "Systematics of the caecilian family Chikilidae (Amphibia: Gymnophiona) with the description of three new species of Chikila from northeast India". Zootaxa. 3666 (4): 401. doi:10.11646/zootaxa.3666.4.1. ISSN 1175-5334.
- Three species of Icthyophis, namely Ichthyophis malabarensis Taylor, 1960, I. peninsularis Taylor, 1960 and I. subterrestris Taylor, 1960, were merged due to lack of phylogenetic deifferentiation vide Gower, D. J.; Dharne, M.; Bhatta, G.; Giri, V.; Vyas, R.; Govindappa, V.; Oommen, O. V.; George, J.; Shouche, Y. & Wilkinson, M. (2007). "Remarkable genetic homogeneity in unstriped, long-tailed Ichthyophis along 1500 km of the Western Ghats, India". Journal of Zoology. 272 (3): 266–275. doi:10.1111/j.1469-7998.2006.00266.x. ISSN 0952-8369.
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