Donax (bivalve)

Species of Donax live, sometimes in high concentrations, vertically aligned in the sand on exposed beaches, on tropical and temperate coasts worldwide. When the waves wash these small clams out of the sand, they can dig back in again quite rapidly. They are filter feeders. Some species, such as Donax variabilis, migrate vertically and horizontally with changes in the tides. These coquina clams are found extensively on the east coast beaches of Trinidad (Mayaro) and widely available in Venezuela. They are called "Chip Chip" in Trinidad & Tobago and "Chipi Chipi" in Venezuela.

Donax cuneatus shell

Species within the genus Donax include:

• Donax asper Hanley 1845[1]
• Donax assimilis Hanley, 1845[1]
• Donax baliregteri Huber, 2012[1]
• Donax bertini Pilsbry, 1901[1]
• Donax bipartitus Sowerby III, 1892[1]
• Donax brazieri Smith, 1892[1]
• Donax bruneirufi Huber, 2012
• Donax burnupi Sowerby III, 1894[1]
• Donax caelatus Carpenter, 1857[1]
• Donax californicus Conrad, 1837[1]
• Donax carinatus Hanley, 1843
• Donax clathratus Reeve, 1855[1]
• Donax culter Hanley, 1845
• Donax cuneatus Linnaeus, 1758[1]
• Donax denticulatus Linnaeus, 1758
• Donax dentifer Hanley, 1843
• Donax ecuadorianus Olsson, 1961[1]
• Donax erythraeensis Bertin, 1881[1]
• Donax faba Gmelin, 1791
• Donax fossor Say, 1822 - was thought at one time to be a northern form of D. variabilis
• Donax francisensis (Cotton & Godfrey, 1938)
• Donax gemmula Morrison, 1971[1]
• Donax gouldii Dall, 1921 - The bean clam, aka Donax gracilis Hanley 1845[1]
• Donax hanleyanus (Philippi, 1847)
• Donax kindermanni (Philippi, 1847)[1]
• Donax lubricus Hanley, 1845[1]
• Donax madagascariensis Wood, 1828
• Donax nitidus Deshayes, 1855[1]
• Donax obesulus Reeve 1854[1]
• Donax obesus d'Orbigny, 1845[1]
• Donax oweni Hanley, 1843
• Donax paxillus Reeve, 1854[1]
• Donax punctatostriatus Hanley, 1843
• Donax rothi Coan, 1983[1]
• Donax semistriatus Poli, 1795[1]
• Donax serra Röding, 1798[1]
• Donax simplex Sowerby III, 1897[1]
• Donax sordidus Hanley, 1845[1]
• Donax striatus Linnaeus, 1767[1]
• Donax texasianus Philippi, 1847[1]
• Donax townsendi Sowerby III, 1894[1]
• Donax transversus Sowerby I, 1825[1]
• Donax trunculus Linnaeus, 1758 - Locally known as tellin in French, tellina or telline in Italian, coquina in Spanish and conquilha or cadelinha in Portuguese.
• Donax variabilis Say, 1822 - Known in Florida under the Spanish word "coquina", is the common and colorful Donax species of the southern Atlantic and Gulf shores from Virginia to the Caribbean. This species is locally abundant on beaches with fine sand from the mid-Atlantic coast to Texas.
• Donax variegatus (Gmelin, 1791)
• Donax vellicatus Reeve, 1855[1]
• Donax veneriformis Lamarck, 1818[1]
• Donax veruinus Hanley, 1913
• Donax vittatus (da Costa, 1778)

The empty small (15 to 25 mm) shells of Donax variabilis and Donax fossor may be found washed up on the beach, especially at low tide. The living animals can often be seen where the waves wash the sand around in the shallowest part of the littoral zone as the tidal level changes. These clams can use the action of waves to move themselves up and down the beach, quickly burrowing into a new location before they can be swept away (the so-called "dance of the coquina").

• Wildlife of Sydney - Fact File - Pipi, Goolwa Cockle, Surf Clam Fact File

• The Nautilus, On the species of Donax of Eastern North America, March 1892, pp. 125-126
• Wayne A. O'Connor & Stephan J. O'Connor, "Early ontogeny of the pipi, Donax (Plebidonax) deltoides (Donacidae; Bivalvia)"; Molluscan Research 31(1): 53–56; ISSN 1323-5818