Sinotyrannus

Sinotyrannus (meaning "Chinese tyrant") is a genus of large basal tyrannosauroid[1] dinosaur, known from a single incomplete fossil specimen including a partial skull, from the Early Cretaceous Jiufotang Formation of Liaoning, China. Specifically, it is a member of the Proceratosauridae, a family that originated in the Jurassic whose members are known from Europe and Asia. Though it is not much younger than primitive tyrannosauroids such as Dilong, it is similar in size to later forms such as Tyrannosaurus. It was much larger than contemporary tyrannosauroids, reaching a total estimated length of 9–10 m (30–33 ft), and was the largest known theropod from the Jiufotang Formation. The type species is S. kazuoensis, described by Ji et al., in 2009.[2]

Sinotyrannus
Temporal range: Early Cretaceous,
Skeletal diagram of Sinotyrannus based on Yutyrannus and Guanlong. Known elements of Sinotyrannus in white (bottom skeleton represents a full skeleton, while the top skeleton represents the holotype).
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Family: †Proceratosauridae
Genus: †Sinotyrannus
Ji et al., 2009
Species:
†S. kazuoensis
Binomial name
†Sinotyrannus kazuoensis
Ji et al., 2009

Description
Estimated size of Sinotyrannus, compared to a human.

Sinotyrannus was a large tyrannosaur, measuring 9–10 metres (30–33 ft) long and weighing 2.5 metric tons (2.8 short tons).[2][3] It was among the largest basal tyrannosauroids known, repudiating the previously presumed trend that tyrannosauroids gradually increased in size throughout the Cretaceous period from small basal forms like Dilong to advanced apex predators such as Tyrannosaurus. The holotype, KZV-001, consists of a disarticulated partial skeleton including the front portion of the skull, three dorsal vertebrae, the incomplete ilia, three articulated manual phalanges (including an ungual), and other fragmentary bones.[2]

The preserved cranial elements include the premaxillae, dentary, and anterior portions of the maxillae and nasals. The dorsal margin of the maxilla is unusually concave unlike the convex condition in tyrannosaurids. The nares are large and elliptical, supporting its relation to proceratosauridae. The dentary gradually curves upwards as it approaches its front edge. Many teeth are preserved attached to the maxillae, with a roughly equal number of denticles on each side, similarly to those of tyrannosaurids. Sinotyrannus could perceivably have had a tall nasal crest like other proceratosaurids, although not enough of its nasals are preserved to be certain.[2][4]

The three preserved vertebrae have very tall neural spines. The proportions of the preserved manual phalanges support the idea that they belong to the second finger, and the ungual has a deep groove on each side. The ilia are mainly present as molds, with the mold of the external side of the left ilium being the most complete. The preacetabular blade is short and wide, with a massive pubic peduncle, while the postacetabular blade is longer and thinner, with a triangular ischial peduncle. These traits of the ilia differentiate it from more advanced tyrannosauroids such as the tyrannosaurids.[2]

Classification
Life reconstruction of S. kazuoensis

The original description of Sinotyrannus proposed that it could have been the earliest tyrannosaurid due to its large size,[2] but subsequent analyses place it as a proceratosaurid tyrannosauroid. It is considered to be part of a clade containing Juratyrant and Stokesosaurus, as they all reputedly share a narrow preacetabular notch.[4] A 2016 analysis instead placed Juratyrant and Stokesosaurus outside of proceratosauridae and proposed that Sinotyrannus is a sister taxon of Yutyrannus within proceratosauridae.[5]

Below is a cladogram by Loewen et al. in 2013.[4]

Tyrannosauroidea
Proceratosauridae

Proceratosaurus bradleyi

Kileskus aristotocus

Guanlong wucaii

Sinotyrannus kazuoensis

Juratyrant langhami

Stokesosaurus clevelandi

Dilong paradoxus

Eotyrannus lengi

Bagaraatan ostromi

Raptorex kriegsteini

Dryptosaurus aquilunguis

Alectrosaurus olseni

Xiongguanlong baimoensis

Appalachiosaurus montgomeriensis

Alioramus altai

Alioramus remotus

Tyrannosauridae

See also

References
  1. Brusatte, S. L.; Norell, M. A.; Carr, T. D.; et al. (2010). "Tyrannosaur paleobiology: new research on ancient exemplar organisms" (PDF). Science. 329 (5998): 1481–1485. Bibcode:2010Sci...329.1481B. doi:10.1126/science.1193304. hdl:20.500.11820/fc52fb23-10e8-466d-a7e9-081260d166c6. PMID 20847260. S2CID 45978858.
  2. Ji, Q.; Ji, S.-A.; Zhang, L.-J. (2009). "First large tyrannosauroid theropod from the Early Cretaceous Jehol Biota in northeastern China" (PDF). Geological Bulletin of China. 28 (10): 1369–1374. Archived from the original (PDF) on 1 October 2018.
  3. Paul, Gregory S. (2010). The Princeton Field Guide to Dinosaurs. New Jersey: Princeton University Press. pp. 99. ISBN 9780691137209.
  4. Loewen, M.A.; Irmis, R.B.; Sertich, J.J.W.; Currie, P. J.; Sampson, S. D. (2013). Evans, David C (ed.). "Tyrant Dinosaur Evolution Tracks the Rise and Fall of Late Cretaceous Oceans". PLoS ONE. 8 (11): e79420. Bibcode:2013PLoSO...879420L. doi:10.1371/journal.pone.0079420. PMC 3819173. PMID 24223179.
  5. Brusatte, Stephen L.; Carr, Thomas D. (2016-02-02). "The phylogeny and evolutionary history of tyrannosauroid dinosaurs". Scientific Reports. 6 (1): 20252. Bibcode:2016NatSR...620252B. doi:10.1038/srep20252. ISSN 2045-2322. PMC 4735739. PMID 26830019.
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