Nacholapithecus
Nacholapithecus kerioi was an ape that lived 14-15 million years ago[1] during the Middle Miocene. Fossils have been found in the Nachola formation in northern Kenya. The only member of the genus Nacholapithecus, it is thought to be a key genus in early hominid evolution. Similar in body plan to Proconsul, it had a long vertebral column with six lumbar vertebrae, no tail, a narrow torso, large upper limbs with mobile shoulder joints, and long feet.[2]
| Nacholapithecus Temporal range:   | |
|---|---|
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| Nacholapithecus kerioi at the Kyoto University Museum | |
| Scientific classification  | |
| Domain: | Eukaryota | 
| Kingdom: | Animalia | 
| Phylum: | Chordata | 
| Class: | Mammalia | 
| Order: | Primates | 
| Suborder: | Haplorhini | 
| Infraorder: | Simiiformes | 
| Family: | †Proconsulidae | 
| Genus: | †Nacholapithecus Ishida, Kunimatsu, Nakatsukasa & Nakano, 1999  | 
| Species | |
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Together with other Kenyapithecinae such as Equatorius, Kenyapithecus, and Griphopithecus, Nacholapithecus displayed synapomorphies with Anoiapithecus.[3]
Taxonomy
    
Nacholapithecus was initially classified as belonging in Kenyapithecus,[4] then attributed[5] to Equatorius (with Equatorius perhaps grouped into a sub-family Equatorinae, instead of both species in Afropithecini),[6][7] finally recognised by Ishida et al. (1999) as a separate genus.[8][9][10] Classified perhaps as a member of the family Proconsulidae.[11]
Fossil finds
    
Nacholapithecus kerioi is known from the lowest part of the Aka Aiteputh Formation, one of five formations in the Neogene System in Nachola, Samburu District, northern Kenya.[12][13] The formation is largely part of the north-western rift flank overlying the Nachola Formation.[14]
Notes
    
- Sawada et al. 2006
 - Henke & Hardt 2007, p. 1020
 - Moyà-Solà et al. 2009
 - Ishida et al. 1984
 - Ward et al. 1999
 - Cameron 2004
 - Cameron 2004, p. 101
 - Ishida et al. 1999
 - Russon & Begun 2004, p. 305
 - Ishida et al. 2004, Abstract
 - Zalmout et al. 2010
 - Nakatsukasa & Kunimatsu 2009, Abstract
 - Sawada et al. 2006
 - Ishida et al. 2006, p. 74
 
References
    
- Cameron, D. W. (2004). Hominid Adaptations and Extinctions. UNSW Press. ISBN 9780868407166.
 - Henke, W.; Hardt, T. (2007). Handbook of paleoanthropology. Springer. ISBN 978-3-540-32474-4.
 - Ishida, H.; Pichford, M.; Nakaya, H.; Nakano, Y. (1984). "Fossil anthropoids from Nachola and Samburu Hills, Samburu District, Kenya". African Study Monographs. Supplementary Issue. 2 (2): 73–85. doi:10.14989/68314.
 - Ishida, H.; Kunimatsu, Y.; Nakatsukasa, M.; Nakano, Y. (1999). "New hominoid genus from the Middle Miocene of Nachola, Kenya". Anthropological Science. 107 (2): 189–191. doi:10.1537/ase.107.189.
 - Ishida, H.; Kunimatsu, Y.; Takano, T.; Nakano, Y.; Nakatsukasa, M. (2004). "Nacholapithecus skeleton from the Middle Miocene of Kenya". Journal of Human Evolution. 46 (1): 69–103. doi:10.1016/j.jhevol.2003.10.001. PMID 14698685.
 - Ishida, H.; Tuttle, R.; Pickford, M.; Ogihara, N.; Nakatsukasa, M. (2006). Ishida, H.; Tuttle, R.; Pickford, M.; Ogihara, N.; Nakatsukasa, M. (eds.). Human Origins and Environmental Backgrounds. Developments in Primatology: Progress and Prospects. Springer. ISBN 9780387296388.
 - Moyà-Solà, S.; Alba, D. M.; Almécija, S.; Casanovas-Vilar, I.; Köhler, M.; de Esteban-Trivigno, S.; Robles, J. M.; Galindo, J.; Fortuny, J. (2009). "A unique Middle Miocene European hominoid and the origins of the great ape and human clade". PNAS. 106 (24): 9601–06. Bibcode:2009PNAS..106.9601M. doi:10.1073/pnas.0811730106. PMC 2701031. PMID 19487676.
 - Nakatsukasa, M.; Kunimatsu, Y. (May–June 2009). "Nacholapithecus and its importance for understanding hominoid evolution". Evolutionary Anthropology. 18 (3): 103–119. doi:10.1002/evan.20208. S2CID 84530387.
 - Russon, A. E.; Begun, D. R. (2004). Russon, A. E.; Begun, D. R. (eds.). The Evolution of Thought: Evolutionary Origins of Great Ape Intelligence. Cambridge University Press. ISBN 9780521783354.
 - Sawada, Y.; Saneyoshi, M.; Nakayama, K.; Sakai, T.; Itaya, T.; Hyodo, M.; Mukokya, Y.; Pickford, M.; Senut, B.; Tanaka, S. (2006). "The Ages and Geological Backgrounds of Miocene Hominoids Nacholapithecus, Samburupithecus, and Orrorin from Kenya". Human Origins and Environmental Backgrounds. Developments in Primatology: Progress and Prospects. Springer. pp. 71–96. doi:10.1007/0-387-29798-7_6. ISBN 978-0-387-29638-8.
 - Ward, S.; Brown, B.; Hill, A.; Kelley, J.; Downs, W. (1999). "Equatorius: A new hominoid genus from the middle Miocene of Kenya". Science. 285 (5432): 1382–1386. doi:10.1126/science.285.5432.1382. PMID 10464093.
 - Zalmout, I. S.; Sanders, W. J.; MacLatchy, L. M.; Gunnell, G. F.; Al-Mufarreh, Y. A.; Ali, M. A.; Nasser, A.-A. H.; Al-Masari, A. M.; Al-Sobhi, S. A.; Nadhra, A. O.; Matari, A. H.; Wilson, J. A.; Gingerich, P. D. (2010). "New Oligocene primate from Saudi Arabia and the divergence of apes and Old World monkeys". Nature. 466 (7304): 360–364. Bibcode:2010Natur.466..360Z. doi:10.1038/nature09094. PMID 20631798. S2CID 205220837.
 
