Lycopsis

Lycopsis; Temporal range: Mid-Late Miocene (Santacrucian-Chasicoan); ~
	Fossil of L. longirostrus
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	†Sparassodonta
Superfamily:	†Borhyaenoidea
Genus:	†Lycopsis; Cabrera 1927
Species
	L. torresi Cabrera 1927 (type species); L. longirostrus Marshall 1976; L. viverensis Forasiepi et al. 2003;
Synonyms
	Anatherium oxyrhynchus Ameghino 1894;

Lycopsis is an extinct genus of South American metatherian, that lived during the Miocene in Argentina and Colombia.[1]

History

Although not named until 1927, Florentino Ameghino described a species now seen as synonymous with Lycopsis torresi, Anatherium oxyrhynchus, in 1895 based on a mandibular ramus with several teeth.[2][3] The fossil was recovered from Puesto Estancia La Costa in Santa Cruz, Argentina, dating to the Miocene.[3] The type material of Lycopsis was collected in July 1895 by "C. Berry" from the middle Miocene strata of the Santa Cruz Formation along the Santa Cruz River in the same area.[2][4] The fossils (MLP 11-113) were fragmentary, constituting only several fragmentary jaw sections from the maxilla and mandible, including several molars.[4][5] However, these fossils were not named until in 1927, Angelo Cabrera named Lycopsis torresi, the generic name meaning "wolf-like aspect" after the anatomy of the mandible and the specific name after Argentine paleontologist and the director of the Museo de la Plata at the time, Luis Maria Torres.[4]

Taxonomy

Skull of Lycopsis

The cladogram after the analysis of Suárez et al., 2015, looks as follows:[6]

Sparassodonta

Patene

Hondadelphys

Stylocynus

Hathliacynidae

Acyon

Cladosictis

Sipalocyon

	Notogale

	Sallacyon

Borhyaenoidea

Lycopsis

Lycopsis longirostrus

Lycopsis viverensis

	Lycopsis torresi

	Lycopsis padillai

	Arctodictis sinclairi

	Arctodictis munizi

Callistoe

Paraborhyaena

Thylacosmilidae

	Patagosmilus

	Thylacosmilus

Distribution

Fossils of Lycopsis have been found in:[3]

Arroyo Chasicó Formation and Santa Cruz Formations, Argentina
Honda Group, Colombia

References

Argot, C. (2004). "Functional-adaptive analysis of the postcranial skeleton of a Laventan borhyaenoid, Lycopsis longirostris (Marsupialia, Mammalia)". Journal of Vertebrate Paleontology. 24 (3): 689–708. doi:10.1671/0272-4634(2004)024[0689:FAOTPS]2.0.CO;2.
Marshall, L. G. (1977). A new species of Lycopsis (Borhyaenidae: Marsupialia) from the La Venta fauna (Late Miocene) of Colombia, South America. Journal of Paleontology, 633-642.
Lycopsis at Fossilworks.org
Cabrera, Á. (1927). Datos para el conocimiento de los dasiuroideos fósiles argentinos. Revista del Museo de la Plata, 30, 271-315.
Argot, C. (2004). Functional-adaptive analysis of the postcranial skeleton of a Laventan borhyaenoid, Lycopsis longirostris (Marsupialia, Mammalia). Journal of Vertebrate Paleontology, 24(3), 689-708.
Suárez, C.; Forasiepi, A. M.; Goin, F. J.; Jaramillo, C. (2015). "Insights into the Neotropics prior to the Great American Biotic Interchange: new evidence of mammalian predators from the Miocene of Northern Colombia". Journal of Vertebrate Paleontology. 36: e1029581. doi:10.1080/02724634.2015.1029581. S2CID 86264178.

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[1] Argot, C. (2004). "Functional-adaptive analysis of the postcranial skeleton of a Laventan borhyaenoid, Lycopsis longirostris (Marsupialia, Mammalia)". Journal of Vertebrate Paleontology. 24 (3): 689–708. doi:10.1671/0272-4634(2004)024[0689:FAOTPS]2.0.CO;2.

[:0-2] Marshall, L. G. (1977). A new species of Lycopsis (Borhyaenidae: Marsupialia) from the La Venta fauna (Late Miocene) of Colombia, South America. Journal of Paleontology, 633-642.

[FWLycopsis-3] Lycopsis at Fossilworks.org

[:1-4] Cabrera, Á. (1927). Datos para el conocimiento de los dasiuroideos fósiles argentinos. Revista del Museo de la Plata, 30, 271-315.

[5] Argot, C. (2004). Functional-adaptive analysis of the postcranial skeleton of a Laventan borhyaenoid, Lycopsis longirostris (Marsupialia, Mammalia). Journal of Vertebrate Paleontology, 24(3), 689-708.

[Suarez-6] Suárez, C.; Forasiepi, A. M.; Goin, F. J.; Jaramillo, C. (2015). "Insights into the Neotropics prior to the Great American Biotic Interchange: new evidence of mammalian predators from the Miocene of Northern Colombia". Journal of Vertebrate Paleontology. 36: e1029581. doi:10.1080/02724634.2015.1029581. S2CID 86264178.