List of recently extinct invertebrates
As of July 2016, the International Union for Conservation of Nature (IUCN) lists 395 extinct species, 206 possibly extinct species, 15 extinct in the wild species, 8 extinct subspecies, and 5 extinct in the wild subspecies of invertebrate.[1][2]
- Extinct in the wild (EW): 15 species
- Critically endangered (CR): 987 species
- Endangered (EN): 1,163 species
- Vulnerable (VU): 2,178 species
- Near threatened (NT, LR/cd): 1,105 species
- Least concern (LC): 6,919 species
- Data deficient (DD): 5,278 species
Invertebrate species (IUCN, 2020.1)
- 17,646 extant species have been evaluated
- 12,368 of those are fully assessed[lower-alpha 1]
- 8024 are not threatened at present[lower-alpha 2]
- 4328 to 9606 are threatened[lower-alpha 3]
- 410 to 616 are extinct or extinct in the wild:
- 395 extinct (EX) species[lower-alpha 4]
- 15 extinct in the wild (EW)
- 206 possibly extinct [CR(PE)]
- 0 possibly extinct in the wild [CR(PEW)]
- excludes data deficient evaluations.
- NT, LR/cd, LC.
- Threatened comprises CR, EN and VU. Upper estimate additionally includes DD.
- Chart omits extinct (EX) species
Nemertea species
Extinct species
- Geonemertes rodericana
Possibly extinct species
- Prosadenoporus agricola
Annelids
Extinct species
- Japan's earthworm (Amynthas japonicus)[3]
- Schmarda's earthworm (Tokea orthostichon)[4][5][6]
- Lake Pedder earthworm (Hypolimnus pedderensis)[7]
Molluscs
Gastropods
Extinct species
- Achatinella abbreviata
- Achatinella buddii
- Achatinella caesia
- Achatinella casta
- Achatinella decora
- Achatinella dimorpha
- Achatinella elegans
- Achatinella juddii
- Achatinella juncea
- Achatinella lehuiensis
- Achatinella livida
- Achatinella papyracea
- Achatinella spaldingi
- Achatinella thaanumi
- Achatinella valida
- Advena campbelli
- Amastra albolabris
- Amastra cornea
- Amastra crassilabrum
- Amastra elongata
- Amastra forbesi
- Amastra pellucida
- Amastra porcus
- Amastra reticulata
- Amastra subrostrata
- Amastra subsoror
- Amastra tenuispira
- Amastra umbilicata
- Amphicyclotulus guadeloupensis
- Shoal sprite (Amphigyra alabamensis)
- Boulder snail (Athearnia crassa)
- Auriculella expansa
- Auriculella uniplicata
- Belgrandiella intermedia
- Bythinella gibbosa
- Bythinella limnopsis
- Bythinella mauritanica
- Bythinella microcochlia
- Bythinella punica
- Caldwellia philyrina
- Campolaemus perexilis
- Carelia anceophila
- Carelia bicolor
- Carelia cochlea
- Carelia cumingiana
- Carelia dolei
- Carelia evelynae
- Carelia glossema
- Carelia hyattiana
- Carelia kalalauensis
- Carelia knudseni
- Carelia lirata
- Carelia lymani
- Carelia mirabilis
- Carelia necra
- Carelia olivacea
- Carelia paradoxa
- Carelia periscelis
- Carelia pilsbryi
- Carelia sinclairi
- Carelia tenebrosa
- Carelia turricula
- Chilonopsis blofeldi
- Chilonopsis exulatus
- Chilonopsis helena
- Chilonopsis melanoides
- Great Saint Helena awl snail (Chilonopsis nonpareil)
- Chilonopsis subplicatus
- Chilonopsis subtruncatus
- Chilonopsis turtoni
- Cahaba pebblesnail (Clappia cahabensis)
- Umbilicate pebblesnail (Clappia umbilicata)
- Collisella edmitchelli
- Colparion madgei
- Ctenoglypta newtoni
- Cyclophorus horridulum
- Cyclosurus mariei
- Mount Matafao different snail (Diastole matafaoi)
- Dupontia proletaria
- Short-spired elimia (Elimia brevis)
- Closed elimia (Elimia clausa)
- Fusiform elimia (Elimia fusiformis)
- Shouldered elimia (Elimia gibbera)
- High-spired elimia (Elimia hartmaniana)
- Constricted elimia (Elimia impressa)
- Hearty elimia (Elimia jonesi)
- Teardrop elimia (Elimia lachryma)
- Ribbed elimia (Elimia laeta)
- Wrinkled elimia (Elimia macglameriana)
- Rough-lined elimia (Elimia pilsbryi)
- Pupa elimia (Elimia pupaeformis)
- Pygmy elimia (Elimia pygmaea)
- Cobble elimia (Elimia vanuxemiana)
- Erepta nevilli
- Fluvidona dulvertonensis
- Gastrocopta chichijimana
- Gastrocopta ogasawarana
- Gibbus lyonetianus
- Gonidomus newtoni
- Gonospira nevilli
- Graecoanatolica macedonica
- Gulella mayottensis
- Excised slitshell (Gyrotoma excisa)
- Striate slitshell (Gyrotoma lewisii)
- Pagoda slitshell (Gyrotoma pagoda)
- Ribbed slitshell (Gyrotoma pumila)
- Pyramid slitshell (Gyrotoma pyramidata)
- Round slitshell (Gyrotoma walkeri)
- Harmogenanina linophora
- Harmogenanina subdetecta
- Helenoconcha leptalea
- Helenoconcha minutissima
- Helenoconcha polyodon
- Helenoconcha pseustes
- Helenoconcha sexdentata
- Helenodiscus bilamellata
- Helenodiscus vernoni
- Heleobia spinellii
- Helicopsis paulhessei
- Hirasea planulata
- Hydrobia gracilis
- Incerticyclus cinereus
- Incerticyclus martinicensis
- Islamia ateni
- Lamellidea monodonta
- Lamellidea nakadai
- Leiorhagium solemi
- Leiostyla lamellosa
- Agate rocksnail (Leptoxis clipeata)
- Oblong rocksnail (Leptoxis compacta)
- Interrupted rocksnail (Leptoxis foremani)
- Maiden rocksnail (Leptoxis formosa)
- Rotund rocksnail (Leptoxis ligata)
- Lyrate rocksnail (Leptoxis lirata)
- Bigmouth rocksnail (Leptoxis occultata)
- Coosa rocksnail (Leptoxis showalterii)
- Squat rocksnail (Leptoxis torrefacta)
- Striped rocksnail (Leptoxis vittata)
- Leucocharis loyaltiensis
- Leucocharis porphyrocheila
- Libera subcavernula
- Libera tumuloides
- Littoraria flammea
- Littoridina gaudichaudii
- Eelgrass limpet (Lottia alveus)
- Lyropupa perlonga
- Olive marstonia (Marstonia olivacea)
- Mautodontha acuticosta
- Mautodontha consimilis
- Mautodontha consobrina
- Mautodontha maupiensis
- Mautodontha parvidens
- Mautodontha punctiperforata
- Mautodontha saintjohni
- Mautodontha subtilis
- Mautodontha unilamellata
- Mautodontha zebrina
- Megalobulimus cardosoi
- Mercuria letourneuxiana
- Nancibella quintalia
- Carinate flat-top snail (Neoplanorbis carinatus)
- Angled flat-top snail (Neoplanorbis smithi)
- Little flat-top snail (Neoplanorbis tantillus)
- Umbilicate flat-top snail (Neoplanorbis umbilicatus)
- Nesopupa turtoni
- Newcombia philippiana
- Ohridohauffenia drimica
- Oleacina guadeloupensis
- Omphalotropis plicosa
- Pachnodus curiosus
- Pachnodus ladiguensis
- Pachnodus velutinus
- Pachystyla rufozonata
- Panulena perrugosa
- Partula approximata
- Partula arguta
- Partula atilis
- Partula aurantia
- Partula auriculata
- Partula bilineata
- Partula callifera
- Partula candida
- Partula castanea
- Partula cedista
- Partula citrina
- Partula compacta
- Partula crassilabris
- Partula cuneata
- Partula cytherea
- Partula dolichostoma
- Partula dolorosa
- Partula eremita
- Partula exigua
- Partula hebe
- Partula filosa
- Partula formosa
- Partula fusca
- Partula garretti
- Polynesia tree snail (Partula imperforata)
- Partula labrusca
- Partula leptochila
- Partula levilineata
- Partula levistriata
- Partula lugubris
- Partula lutea
- Partula microstoma
- Partula navigatoria
- Partula ovalis
- Partula planilabrum
- Partula producta
- Partula protea
- Partula protracta
- Partula radiata
- Partula raiatensis
- Partula remota
- Partula robusta
- Partula rustica
- Partula sagitta
- Mount Alifana partula (Partula salifana)
- Partula thalia
- Partula turgida
- Partula umbilicata
- Partula variabilis
- Partula vittata
- Partulina crassa
- Partulina montagui
- Perdicella fulgurans
- Perdicella maniensis
- Perdicella zebra
- Perdicella zebrina
- Fish Lake physa (Physella microstriata)
- Placostylus cuniculinsulae
- Acorn ramshorn (Planorbella multivolvis)
- Plectostoma sciaphilum
- Pleurodonte desidens
- Posticobia norfolkensis
- Pseudamnicola barratei
- Pseudamnicola desertorum
- Pseudamnicola doumeti
- Pseudamnicola globulina
- Pseudamnicola latasteana
- Pseudamnicola oudrefica
- Pseudamnicola ragia
- Pseudamnicola singularis
- Pseudocampylaea loweii
- Pseudohelenoconcha spurca
- Pupilla obliquicosta
- Corded purg (Pyrgulopsis nevadensis)
- Quintalia flosculus
- Quintalia stoddartii
- Rhachis comorensis
- Rhachis sanguineus
- Aldabra banded snail (Rhachistia aldabrae)
- Samoana inflata
- Samoana jackieburchi
- Sinployea canalis
- Sinployea decorticata
- Sinployea harveyensis
- Sinployea otareae
- Sinployea planospira
- Sinployea proxima
- Sinployea rudis
- Sinployea tenuicostata
- Sinployea youngi
- Ouachita pebblesnail (Somatogyrus amnicoloides)
- Thick-lipped pebblesnail (Somatogyrus crassilabris)
- Channeled pebblesnail (Somatogyrus wheeleri)
- Fish springs marshsnail (Stagnicola pilsbryi)
- Taipidon anceyana
- Taipidon marquesana
- Taipidon octolamellata
- Thaumatodon multilamellata
- Tomigerus gibberulus
- Tomigerus turbinatus
- Tornelasmias capricorni
- Trochoidea picardi
- Tropidophora desmazuresi
- Tropidophora semilineata
- Vitrinula chaunax
- Vitrinula chichijimana
- Vitrinula hahajimana
Possibly extinct species
- Aaadonta angaurana
- Aaadonta kinlochi
- Aaadonta pelewana
- Alzoniella galaica
- Beddomeia tumida
- Belgrandia moitessieri
- Belgrandia varica
- Belgrandiella boetersi
- Belgrandiella cavernica
- Belgrandiella kreisslorum
- Belgrandiella multiformis
- Bulimulus achatellinus
- Bulimulus adelphus
- Bulimulus deridderi
- Bulimulus duncanus
- Bulimulus eos
- Bulimulus lycodus
- Bulimulus saeronius
- Bulimulus sp. nov. 'josevillani'
- Bulimulus sp. nov. 'krameri'
- Bulimulus sp. nov. 'nilsodhneri'
- Bulimulus sp. nov. 'tuideroyi'
- Bulimulus sp. nov. 'vanmoli'
- Bulimulus tanneri
- Bythinella eutrepha
- Bythiospeum dubium
- Bythiospeum gonostoma
- Bythiospeum putei
- Bythiospeum turritum
- Coneuplecta turrita
- Conturbatia crenata
- Delos gardineri
- Dianella schlickumi
- Diplommatina alata
- Diplommatina aurea
- Diplommatina gibboni
- Discula lyelliana
- Discula tetrica
- Mossy elimia (Elimia troostiana)
- Endodonta apiculata
- Eua globosa
- Falsipyrgula beysehirana
- Gabbiella matadina
- Geomitra delphinuloides
- Geomitra grabhami
- Graecoanatolica brevis
- Graecoanatolica conica
- Hemicycla modesta
- Henrigirardia wienini
- Hydrobia anatolica
- Iglica gratulabunda
- Islamia bendidis
- Islamia graeca
- Islamia hadei
- Islamia pseudorientalica
- Islamia zermanica
- Kirelia carinata
- Kubaryia pilikia
- Lauopa mbalavuana
- Leiostyla abbreviata
- Leiostyla cassida
- Leiostyla gibba
- Leiostyla simulator
- Madgeaconcha sevathiani
- Margarya yangtsunghaiensis
- Ozark pyrg (Marstonia ozarkensis)
- Melanoides agglutinans
- Melanopsis germaini
- Melanopsis infracincta
- Melanopsis khabourensis
- Melanopsis pachya
- Mercuria punica
- Monilearia pulverulenta
- Montserratina becasis
- Neritina tiassalensis
- Ohridohauffenia minuta
- Omphalotropis ingens
- Opisthostoma decrespignyi
- Opisthostoma otostoma
- Paladilhiopsis janinensis
- Palaina albata
- Palaina patula
- Palaina platycheilus
- Palaina pupa
- Partula guamensis
- Partula leucothoe
- Partula milleri
- Placostylus koroensis
- Plectostoma charasense
- Plectostoma dindingensis
- Plectostoma turriforme
- Potamopyrgus acus
- Pseudogibbula cara
- Samoana cramptoni
- Sardohoratia sulcata
- Soapitia dageti
- Stocky pebblesnail (Somatogyrus crassus)
- Tennessee pebblesnail (Somatogyrus currierianus)
- Fluted pebblesnail (Somatogyrus hendersoni)
- Atlas pebblesnail (Somatogyrus humerosus)
- Dwarf pebblesnail (Somatogyrus nanus)
- Spiralix corsica
- Thickshell pondsnail (Stagnicola utahensis)
- Succinea rotumana
- Tanousia zrmanjae
- Brune's tryonia (Tryonia brunei)
- Valvatorbis mauritii
- Zaumia sanctizaumi
- Zingis radiolata
Extinct in the wild species
Extinct subspecies
- Achatinella apexfulva vittata
- Achatinella bulimoides rosea
- Partula suturalis suturalis
- Partula taeniata taeniata
- Placostylus bivaricosus etheridgei
Extinct in the wild subspecies
- Partula suturalis strigosa
- Partula suturalis vexillum
- Partula taeniata elongata
- Partula taeniata nucleola
- Partula taeniata simulans
Bivalvia
Extinct species
- Coosa elktoe (Alasmidonta mccordi)
- Carolina elktoe (Alasmidonta robusta)
- Ochlockonee arcmussel (Alasmidonta wrightiana)
- Chambardia letourneuxi
- Arc-form pearly mussel (Epioblasma arcaeformis)
- Angled riffleshell (Epioblasma biemarginata)
- Arcuate pearly mussel (Epioblasma flexuosa)
- Acorn pearly mussel (Epioblasma haysiana)
- Narrow catspaw (Epioblasma lenior)
- Forkshell (Epioblasma lewisii)
- Fine-rayed pearly mussel (Epioblasma personata)
- Nearby pearly mussel (Epioblasma propinqua)
- Sampson's naiad (Epioblasma sampsonii)
- Cumberland leafshell (Epioblasma stewardsonii)
- Turgid riffle shell (Epioblasma turgidula)
- Lined pocketbook (Lampsilis binominata)
- Tombigbee moccasinshell (Medionidus mcglameriae)
- Highnut (Pleurobema altum)
- Hazel pigtoe (Pleurobema avellanum)
- Scioto pigtoe (Pleurobema bournianum)
- Yellow pigtoe (Pleurobema flavidulum)
- Brown pigtoe (Pleurobema hagleri)
- Alabama pigtoe (Pleurobema johannis)
- Coosa pigtoe (Pleurobema murrayense)
- Longnut (Pleurobema nucleopsis)
- Ovate clubshell (Pleurobema perovatum)
- Alabama clubshell (Pleurobema troschelianum)
- True pigtoe (Pleurobema verum)
- Unio cariei
Possibly extinct species
- Cuneopsis demangei
- Dreissena caspia
- Southern acorn riffle shell (Epioblasma othcaloogensis)
- Lamprotula crassa
- Lamprotula liedtkei
- Lamprotula nodulosa
- Haddleton lampmussel (Obovaria haddletoni)
- Black clubshell (Pleurobema curtum)
Extinct subspecies
- Yellow-blossom (Epioblasma florentina florentina)
- Epioblasma torulosa gubernaculum
- Tubercled-blossom pearly mussel (Epioblasma torulosa torulosa)
Cnidaria
Possibly extinct species
- Millepora boschmai
- Wellington's solitary coral (Rhizopsammia wellingtoni)
Arthropods
Arachnids
Extinct species
- Centrobunus braueri
- Gardiner's giant mite (Dicrogonatus gardineri)
- Hirstienus nanus
- Metazalmoxis ferruginea
- Peromona erinacea
- Pleorotus braueri
- Sitalcicus gardineri
- Stipax triangulifer
- Thomasettia seychellana
Possibly extinct species
- Afrogarypus seychellesensis
- Biantes parvulus
- Euso muehlenbergi
- Gamasomorpha austera
- Holozoster ovalis
- Ibalonius lomani
- Idioctis intertidalis
- Mitraceras crassipalpum
- Moneta coercervea
- Nesiergus gardineri
- Nesiergus halophilus
- Paccius quadridentatus
- Seychellia lodoiceae
- Sitalcicus incertus
- Steriphopus lacertosus
- Voraptus tenellus
Millipedes
Extinct species
- Eucarlia alluaudi
- Orthomorpha crinita
- Spirobolellus praslinus
Possibly extinct species
- Diglossosternoides curiosus
- Rhinotus albifrons
- Spirobolellus simplex
Malacostracans
Includes crabs, lobsters, crayfish, shrimp, krill, woodlice, and many others.
Extinct species
- Cambarellus alvarezi
- Macrobrachium leptodactylus
- Sooty crayfish (Pacifastacus nigrescens)
- Sandhills crayfish (Procambarus angustatus)
- Rubious cave amphipod (Stygobromus lucifugus)
- Pasadena freshwater shrimp (Syncaris pasadenae)
Possibly extinct species
- Atya brachyrhinus
- Cambarellus areolatus
- White spring cave crayfish (Cambarus veitchorum)
- Caridina apodosis
- Caridina yilong
- Cryphiops luscus
- Macrobrachium denticulatum
- Macrobrachium oxyphilus
- Macrobrachium purpureamanus
- Macrobrachium scorteccii
- Florida cave shrimp (Palaemonetes cummingi)
- Perbrinckia gabadagei
- Big-cheeked cave crayfish (Procambarus delicatus)
- Procambarus paradoxus
- Sinodina acutipoda
- Strengeriana antioquensis
- Tehuana veracruzana
Extinct in the wild species
- Socorro isopod (Thermosphaeroma thermophilum)
Insects
Extinct species
- Pecatonica river mayfly (Acanthametropus pecatonica)
- Poko noctuid moth (Agrotis crinigera)
- Midway noctuid moth (Agrotis fasciata)
- Kerr's noctuid moth (Agrotis kerri)
- Laysan noctuid moth (Agrotis laysanensis)
- Agrotis photophila
- Procellaris grotis noctuid moth (Agrotis procellaris)
- Robert's stonefly (Alloperla roberti)
- Chestnut ermine moth (Argyresthia castaneela)
- Campsicnemus mirabilis
- Clavicoccus erinaceus
- Coleophora leucochrysella
- Central Valley grasshopper (Conozoa hyalina)
- Deloneura immaculata
- Drosophila lanaiensis
- Dryophthorus distinguendus
- American chestnut moth (Ectodemia castaneae)
- Phleophagan chestnut moth (Ectodemia phleophaga)
- Genophantis leahi
- Xerces blue (Glaucopsyche xerces)
- Confused moth (Helicoverpa confusa)
- Minute noctuid moth (Helicoverpa minuta)
- Tobias' caddisfly (Hydropsyche tobiasi)
- Mono Lake diving beetle (Hygrotus artus)
- Laysan dropseed noctuid moth (Hypena laysanensis)
- Hilo noctuid moth (Hypena newelli)
- Lovegrass noctuid moth (Hypena plagiota)
- Kaholuamano noctuid moth (Hypena senicula)
- Saint Helena earwig (Labidura herculeana)
- Lepidochrysops hypopolia
- Levuana moth (Levuana iridescens)
- Libythea cinyras
- Margatteoidea amoena
- Mecodema punctellum
- Megadytes ducalis
- Maui upland damselfly (Megalagrion jugorum)
- Rocky Mountain locust (Melanoplus spretus)
- Antioch dunes shieldback katydid (Neduba extincta)
- Oeobia sp. nov.
- Laysan weevil (Oodemas laysanensis)
- Robust burrowing mayfly (Pentagenia robusta)
- Phyllococcus oahuensis
- Ridley's stick insect (Pseudobactricia ridleyi)
- Rhantus novacaledoniae
- Rhantus orbignyi
- Rhantus papuanus
- Castle Lake caddisfly (Rhyacophila amabilis)
- Rhyncogonus bryani
- Kona giant looper moth (Scotorythra megalophylla)
- Ko'olau giant looper moth (Scotorythra nesiotes)
- Perrin's cave beetle (Siettitia balsetensis)
- Volutine stoneyian tabanid fly (Stonemyia volutina)
- Chestnut clearwing moth (Tischeria perplexa)
- Athens caddisfly (Triaenodes phalacris)
- Three-tooth caddisfly (Triaenodes tridonata)
- Fort ross weevil (Trigonoscuta rossi)
- Yorba linda weevil (Trigonoscuta yorbalindae)
- Ola'a peppered looper moth (Tritocleis microphylla)
Possibly extinct species
- Morogoro pretty grasshopper (Acanthothericles bicoloripes)
- Zanzibar giant forest grasshopper (Allaga ambigua)
- Spined dwarf mantis (Ameles fasciipennis)
- Andrena labiatula
- Usambara splendid grasshopper (Anischnansis burtti)
- Anisogomphus solitaris
- Balta crassivenosa
- Bombus rubriventris
- Togo red jewel (Chlorocypha jejuna)
- Morogoro monkey grasshopper (Chromomastax movogovodia)
- Mpwapwa silent grasshopper (Chromousambilla burtti)
- Maspalomas bow-legged grasshopper (Dericorys minutus)
- Disparoneura ramajana
- Drepanosticta adami
- Drepanosticta austeni
- Drepanosticta montana
- Drepanosticta submontana
- Enallagma maldivensis
- Kilosa noble grasshopper (Eupropacris abbreviata)
- Mlingano monkey grasshopper (Euschmidtia bidens)
- Burtt's monkey grasshopper (Euschmidtia burtti)
- Dirsh's monkey grasshopper (Euschmidtia dirshi)
- Phipps' monkey grasshopper (Euschmidtia phippsi)
- Dar-es-salaam monkey grasshopper (Euschmidtia viridifasciata)
- Gran Canaria bush-cricket (Evergoderes cabrerai)
- Heliogomphus lyratus
- Heliogomphus nietneri
- Heteragrion peregrinum
- Holocompsa pusilla
- Macromia flinti
- Megachile cypricola
- Molokai damselfly (Megalagrion molokaiense)
- Metaleptobasis gibbosa
- Nomada siciliensis
- Elusive skimmer (Orthetrum rubens)
- Palaemnema edmondi
- Zulu ambush katydid (Peringueyella zulu)
- Perissolestes remus
- Pieris wollastoni
- Seychelles shortwinged groundhopper (Procytettix fusiformis)
- Three-lobed bush-cricket (Rhacocleis trilobata)
- Sliferia similis
- Theganopteryx grisea
- Theganopteryx liturata
- Theganopteryx scotti
- Peringuey's seedpod shieldback (Thoracistus peringueyi)
Extinct in the wild species
- Oahu deceptor bush cricket (Leptogryllus deceptor)
See also
References
- "IUCN Red List version 2016.1". The IUCN Red List of Threatened Species. International Union for Conservation of Nature and Natural Resources (IUCN). Retrieved 4 July 2016.
- "Table 9: Possibly Extinct and Possibly Extinct in the Wild Species (IUCN Red List version 2016-1)". The IUCN Red List of Threatened Species. Union for Conservation of Nature and Natural Resources (IUCN). Retrieved 4 July 2016.
- Blakemore (2019). "Extinction of Japan's first formally described earthworm Amynthas japonicus (Horst, 1883) (Annelida, Oligochaeta, Megadrilacea, Megascolecidae)" (PDF). Bulletin of Kanagawa Museum. 48: 55–60. Retrieved 10 May 2019.
- Blakemore (2012). "On Schmarda's lost earthworm and some newly found New Zealand species (Oligochaeta: Megadrilacea: Lumbricidae, Acanthodrilidae, Octochaetidae, & Megascolecidae s. stricto)". Journal of Species Research. JSR. 1 (2): 105–132. doi:10.12651/jsr.2012.1.2.105.
- Blakemore, R.J. (2017). "Tokea orthostichon". IUCN Red List of Threatened Species. 2017: e.T103192401A103193535. doi:10.2305/IUCN.UK.2017-2.RLTS.T103192401A103193535.en. Retrieved 16 October 2022.
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