Dryosauridae

Dryosauridae; Temporal range: Middle Jurassic – Early Cretaceous,
	Dysalotosaurus skeletal mount in Berlin
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Clade:	†Ornithischia
Clade:	†Ornithopoda
Clade:	†Iguanodontia
Clade:	†Dryomorpha
Superfamily:	†Dryosauroidea; Milner & Norman, 1984
Family:	†Dryosauridae; Milner & Norman, 1984
Subgroups
	†Callovosaurus; †Dryosaurus; †Dysalotosaurus; †Elrhazosaurus; †Eousdryosaurus?; †Iyuku; †Kangnasaurus?; †Valdosaurus;

Dryosauridae was a family of primitive iguanodonts, first proposed by Milner & Norman in 1984. They are known from Middle Jurassic to Early Cretaceous rocks of Africa, Europe, and North America.[1][2][3][4]

Phylogeny

Until recently many dryosaurids have been regarded as dubious (Callovosaurus and Kangnasaurus) or as species of the type member, Dryosaurus (Dysalotosaurus, Elrhazosaurus and Valdosaurus). However, more recent studies redescribe these genera as valid.[3][4][5][6][7] The cladogram below follows Paul M. Barrett, Richard J. Butler, Richard J. Twitchett and Stephen Hutt (2011).[7]

Rhabdodontidae

	Muttaburrasaurus

	Rhabdodon

	Zalmoxes

Dryosauridae

	Valdosaurus

	Elrhazosaurus

References

Norman, David B.; Weishampel, David B. (1990). "Iguanodontidae and related ornithopods". In Weishampel, David B.; Dodson, Peter; Osmólska Halszka (eds.). The Dinosauria. Berkeley: University of California Press. pp. 510–533. ISBN 0-520-06727-4.
Norman, David B. (2004). "Basal Iguanodontia". In Weishampel, D.B.; Dodson, P.; Osmólska H. (eds.). The Dinosauria (2nd ed.). Berkeley: University of California Press. pp. 413–437. ISBN 0-520-24209-2.
Ruiz-Omeñaca, José Ignacio; Pereda Suberbiola Xabier; Galton, Peter M. (2007). "Callovosaurus leedsi, the earliest dryosaurid dinosaur (Ornithischia: Euornithopoda) from the Middle Jurassic of England". In Carpenter Kenneth (ed.). Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. Bloomington and Indianapolis: Indiana University Press. pp. 3–16. ISBN 978-0-253-34817-3.
Galton, Peter M. (2009). "Notes on Neocomian (Lower Cretaceous) ornithopod dinosaurs from England – Hypsilophodon, Valdosaurus, "Camptosaurus", "Iguanodon" – and referred specimens from Romania and elsewhere" (PDF). Revue de Paléobiologie. 28 (1): 211–273.
McDonald, A.T.; Kirkland, J.I.; DeBlieux, D.D.; Madsen, S.K.; Cavin, J.; Milner, A.R.C.; Panzarin, L. (2010). "New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs". PLOS ONE. 5 (11): e14075. doi:10.1371/journal.pone.0014075. PMC 2989904. PMID 21124919.
Andrew T. McDonald (2011). "The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)" (PDF). Zootaxa. 2783: 52–68. doi:10.11646/zootaxa.2783.1.4.
Paul M. Barrett; Richard J. Butler; Richard J. Twitchett; Stephen Hutt (2011). "New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of southern England". Special Papers in Palaeontology. 86: 131–163.

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[NW90-1] Norman, David B.; Weishampel, David B. (1990). "Iguanodontidae and related ornithopods". In Weishampel, David B.; Dodson, Peter; Osmólska Halszka (eds.). The Dinosauria. Berkeley: University of California Press. pp. 510–533. ISBN 0-520-06727-4.

[DBN04-2] Norman, David B. (2004). "Basal Iguanodontia". In Weishampel, D.B.; Dodson, P.; Osmólska H. (eds.). The Dinosauria (2nd ed.). Berkeley: University of California Press. pp. 413–437. ISBN 0-520-24209-2.

[JROetal07-3] Ruiz-Omeñaca, José Ignacio; Pereda Suberbiola Xabier; Galton, Peter M. (2007). "Callovosaurus leedsi, the earliest dryosaurid dinosaur (Ornithischia: Euornithopoda) from the Middle Jurassic of England". In Carpenter Kenneth (ed.). Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. Bloomington and Indianapolis: Indiana University Press. pp. 3–16. ISBN 978-0-253-34817-3.

[PMG09-4] Galton, Peter M. (2009). "Notes on Neocomian (Lower Cretaceous) ornithopod dinosaurs from England – Hypsilophodon, Valdosaurus, "Camptosaurus", "Iguanodon" – and referred specimens from Romania and elsewhere" (PDF). Revue de Paléobiologie. 28 (1): 211–273.

[iguanacolossus-5] McDonald, A.T.; Kirkland, J.I.; DeBlieux, D.D.; Madsen, S.K.; Cavin, J.; Milner, A.R.C.; Panzarin, L. (2010). "New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs". PLOS ONE. 5 (11): e14075. doi:10.1371/journal.pone.0014075. PMC 2989904. PMID 21124919.

[McDonald-6] Andrew T. McDonald (2011). "The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)" (PDF). Zootaxa. 2783: 52–68. doi:10.11646/zootaxa.2783.1.4.

[Phylogeny-7] Paul M. Barrett; Richard J. Butler; Richard J. Twitchett; Stephen Hutt (2011). "New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of southern England". Special Papers in Palaeontology. 86: 131–163.