Xenophoridae

The Xenophorids live on sand and mud bottoms of the continental shelves and the continental slopes of the subtropical and tropical seas and range from very shallow water to depths of more than 1,400 meters.[1]

Xenophorids are unusual in that in many of the species the animal cements small stones or shells to the edge of the shell as it grows, thus the shells of those species are sometimes humorously referred to as "shell-collecting shells". The genus name Xenophora comes from two ancient Greek words and means "bearing (or carrying) foreigners". The shells are small to rather large (diameter of base without attachments 19–160 mm; height of shell 21–100 mm), depressed to conical, with narrow to wide, simple to spinose peripheral edge or flange separating spire from base. Aperture large, base broad, rather flattened, often umbilicate. Periostracum very thin or wanting. Protoconch depressed-conical, multispiral (in one species paucispiral). Teleoconch usually with foreign objects attached in spiral series to peripheral flange and, sometimes, remainder of dorsum, at least on early whorls. Operculum horny, yellowish to brown, nucleus lateral, with simple growth lamellae, sometimes with conspicuous radial striae or hollow radial ribs.[1]

Aperture view of a xenophorid gastropod (Xenophora pallidula).

Genera within the family Xenophoridae include:[1][2]

• † Acanthoxenophora Perrilliat & Vega, 2001
• Aspidophoreas Nappo, Bini & Santucci, 2022
• Onustus Swainson, 1840 - synonyms: Trochotugurium Sacco, 1896; Tugurium Fischer in Kiener, 1879[3]
• Ponderiana Nappo, Bini & Santucci, 2022
• Stellaria Möller, 1832 - synonym: Haliphoebus Fischer in Kiener, 1879; Xenophora (Stellaria) Schmidt, 1832[4]
• Xenophora Fischer von Waldheim, 1807[5] - type genus

• Kreipl K. & Alf A. (1999). Recent Xenophoridae. Conchbooks, Hackenheim > Germany. 148pp.
• Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526

Wikimedia Commons has media related to Xenophoridae.

• Family: Xenophoridae (Carrier Shells)
• Troschel, F. H. (1852). Bericht über die Leistungen im Gebiete in der Naturgeschichte der Mollusken während des Jahres 1851. Archiv für Naturgeschichte. 18(2): 257-307
• Gray, J. E. (1840). Shells of molluscous animals, pp. 105-152. in: Synopsis of the contents of the British Museum. ed. 42. G. Woodfall. London. 370 pp.
• Adams, H. & Adams, A. (1853-1858). The genera of Recent Mollusca; arranged according to their organization. London, van Voorst. Vol. 1: xl + 484 pp.; vol. 2: 661 pp.; vol. 3: 138 pls.
• Ladd, H.S. (1977). Cenozoic fossil mollusks from western Pacific islands; Gastropods (Eratoidae through Harpidae). U.S. Geological Survey Professional Paper. 533: i–iv, 1–84, pls 1–23 This article incorporates text from this source, which is in the public domain.
• Ponder W.F. (1983). A revision of the Recent Xenophoridae of the World and of the Australian Fossil Species (Mollusca : Gastropoda). Memoir 17. The Australian Museum Sydney, Australia
• Irwin, A. R.; Strong, E. E.; Kano, Y.; Harper, E. M.; Williams, S. T. (2021). Eight new mitogenomes clarify the phylogenetic relationships of Stromboidea within the caenogastropod phylogenetic framework. Molecular Phylogenetics and Evolution. 158: 107081